Natural History of the Mammalia of India and Ceylon

NO. 427. EQUUS HEMIONUSThe Kiang or Wild Ass of Thibet

NATIVE NAMES.—Kiang or Dizightai, Thibetan.

HABITAT.—Thibet and Central Asia; Ladakh.

DESCRIPTION.—Darker in hue than the ghor-khur, especially on the flanks, contrasting abruptly with the white of the under-parts. It has the dark line along the back, but not the cross band on the shoulder; ears shorter.

SIZE.—About 12 to 14 hands in height.

From its larger size, shorter ears, and its shrill bray, which has been mistaken for a neigh, this animal has at times been taken for a horse, and described as such. The kiang, of which there is a living specimen in the London Zoological Gardens, inhabits the high plateaux of Thibet, ranging up to fifteen and sixteen thousand feet above the sea level. It is very swift and wary.

The late Brigadier-General McMaster, in his 'Notes on Jerdon,' page 248, says: "An excellent sportsman and very close observer, who, being a cavalry officer, should be able to give a sound opinion on the matter, assured me that the voice of the wild horse of the snowy Himalayas is 'an unmistakeable neigh, not a bray,' and that he certainly looked on them as horses. He had seen several of these animals, and killed one." Captain (now General) R. Strachey wrote of it: "My impression as to the voice of the kyang is that it is a shrieking bray and not a neigh;" and again: "the kyang, so far as external aspect is concerned, is obviously an ass and not an horse." Of this there is but little doubt. Moorcroft, in his travels, vol. i. p. 312, states: "In the eastern parts of Ladakh is a nondescript wild variety of horse which I may call Equus kiang. It is perhaps more of an ass than a horse, but its ears are shorter, and it is certainly not the gur-khor or wild ass of Sind." Further on, at page 442, he-adds: "We saw many herds of the kyang, and I made numerous attempts to bring one down, but with invariably bad success. Some were wounded, but not sufficiently to check their speed, and they quickly bounded up the rocks, where it was impossible to follow. They would afford excellent sport to four or five men well mounted, but a single individual has no chance. The kyang allows his pursuer to approach no nearer than five or six hundred yards; he then trots off, turns, looks and waits till you are almost within distance, when he is off again. If fired at he is frightened, and scampers off altogether. The Chanthan people sometimes catch them by snares—sometimes shoot them. From all I have seen of the animal I should pronounce him to be neither a horse nor an ass. His shape is as much like that of the one as the other, but his cry is more like braying than neighing. The prevailing colour is a light reddish-chestnut, but the nose, the under-part of the jaw and neck, the belly and the legs are white, the mane is dun and erect, the ears are moderately long, the tail bare and reaching a little below the hock. The height is about fourteen hands. The form, from the fore to the hind leg and feet to a level with the back is more square than that of an ass. His back is less straight, and there is a dip behind the withers and a rounding of the crupper which is more like the shape of the horse; his neck also is more erect and arched than that of the ass. He is perhaps more allied to the quagga, but without stripes, except a reported one along each side of the back to the tail. These were seen distinctly in a foal, but were not distinguished in the adults."

FAMILY TAPIRIDÆ—THE TAPIRS

These are somewhat hog-like animals, with elongated snouts, possessing four toes on their fore-feet, and three on the hinder ones. They live in dense forests, are nocturnal in habit, and live exclusively on a vegetable diet. The Indian tapir has a more powerful and extensile trunk than the American, and its skull shows in consequence a greater space for the attachment of the muscles. The dentition is as follows:—Inc., 3—3/3—3; can., 1—1/1—1; premolars, 4—4/4—4; molars, 3—3/3—3. The outer incisors somewhat resemble canines, whilst the others are very small. The canines themselves are not large.

The tapir is not found in India proper, but the Malayan species is occasionally to be come across in Burmah, having been killed in Tenasserim.

GENUS TAPIRUSNO. 428. TAPIRUS MALAYANUSThe Malay Tapir

NATIVE NAMES.—Ta-ra-shu, Burmese; Kuda-ayer, Malayan; Sala-dang of the Limuns in Sumatra; Gindol of the Mannas in Sumatra; Babi-alu in Bencoolen; Tennu in Malacca.

HABITAT.—Tenasserim provinces, as high as the fifteenth degree north latitude; Lower Siam; the Malayan peninsula; Sumatra and Borneo.

DESCRIPTION.—General colour glossy black, but with the back, rump, and sides of the belly white. The young are beautifully variegated, being striped and spotted with yellow fawn on the upper parts of the body, and with white below.

Mr. Mason writes: "Though seen so rarely, the tapir is by no means uncommon in the interior of the Tavoy and Mergui provinces. I have frequently come upon its recent footmarks, but it avoids the inhabited parts of the country. It has never been heard of north of the valley of the Tavoy river."

The tapir is naturally all the world over a very shy, retiring animal, but it is capable of being tamed when taken young, and of showing great attachment.

FAMILY RHINOCEROTIDÆ

"The skeleton of the rhinoceros viewed generally has a resemblance to that of the little hyrax, the tapir, and the horse. The skull is very much elevated at the base, being somewhat of a pyramidal form, and the nasal bones curve upwards and downwards, and are of such a size and thickness, in order to support one or more immense horns, that they are quite unparalleled for their development in any other existing quadruped. The nasal bones, together with the premaxillary and maxillary bones, form the general contour for the external apertures of the nostrils. This is peculiar, and found in no other animal with the exception of the tapir."—Prof. W. Boyd Dawkins and Mr. Oakley.

The external appearance of this animal is familiar to most—a large ungainly creature, with a long head, a massive horn on its nose, sometimes two horns; a round unwieldly body covered with an immensely thick hide arranged in heavy folds; short tail and short legs, with three toes covered with broad nails or hoofs.

The stomach is simple; the intestines about eight times the length of the body, and the cæcum is large and sacculated. The horn is a mere agglutinated mass of hair or fibre superimposed on the skin, and has no bony core. The females have two inguinal mammæ.

The dentition is peculiar; "the grinders are implanted by distinct roots, and in the upper jaw their crowns are traversed by two deep folds of enamel which constitute open valleys. In the lower jaw they are composed of two crescent-shaped lobes, also open. The covering of cement is thin, and never fills up the valleys, as in the case of the more complex dental system in the horse. The normal number of grinders is seven in each jaw, while the incisors, as we have already remarked, vary not only in form but also are sometimes absent, and canines are not developed in any of the living or fossil members of the family."—Boyd Dawkins and Oakley.

The Rhinocerotidæ are divided into two groups—the Asiatic and the African; and the former consist of two genera—RHINOCEROS and CERATORHINUS, the former with one and the latter with two horns.

It is a moot point whether the rhinoceros is or is not the unicorn of Scripture, though it is by no means clear that the animal in question was a one-horned creature, but according to some might have been the great wild ox or urus of Macedonia. An Indian single-horned rhinoceros was sent from India to the king of Portugal in 1513, and from it various most distorted pictures were disseminated throughout Europe. It was represented as covered with a wondrous suit of armour beautifully decorated, and with a second horn on its shoulders!

The first one brought alive to England was in 1685. Parsons describes and figures one brought to Europe in 1739, and another in 1741 ('Philosophical Transactions,' xlii.).

The Asiatic rhinoceroses differ from the African in having the skin divided into shields by well-marked folds, long upper cutting teeth, the African having none, and by the produced conical nasal bones of the skull instead of broad and rounded ones. There are one or two other minor yet well-marked differences which we need not mention here.

GENUS RHINOCEROS

"The skin divided into shields by well-marked folds, lumbar and neck-folds well developed; horn single, anterior; part of occipital bone near the occipital condyle and the condyles themselves prominent."—Gray.

There are two species in India, viz. Rhinoceros Indicus and R. Sondaicus, the latter being the Javan species.

For the following description of the former I have to thank Mr. J. Cockburn, who, with most unselfish kindness, kept back the article he was about to publish, and gave it to me to incorporate in this work. The following remarks on dentition are also his:30—

"The normal dentition of R. Indicus is: Inc., 1—1/2—2; premolars, 4—4/4—4; molars, 3—3/3—3; but the dentition varies to a great extent; for example, in a specimen of R. Sondaicus it stood: Inc., 1—1/2—2; molars, 6—7/6—6. The first premolar in both Indicus and Sondaicus is a deciduous tooth, which is not usually replaced, and gradually drops out with age, but it may be retained till extreme old age. In the majority of cases it is either lost or worn down before the last molar is in wear. The incisors also vary greatly in the adult animal; they are 1—1/2—2, the outer pair below being the formidable dagger-shaped tushes, with which they inflict the terrible gashes they can produce. The median pair lower are usually lost or absorbed by advancing age, having no functions, and the incisive tusks themselves are subject to very rapid wear, being often worn down before the animal has reached middle age. Occasionally R. Indicus has six incisors in the lower jaw (the normal number in other mammalia), and four in the upper, but this is very exceptional."—J. Cockburn, MS.

NO. 429. RHINOCEROS INDICUS(Jerdon's No. 212)

NATIVE NAMES.—Genda, Gonda, Ganda, or Genra, Hindi; Gor, Assamese.

HABITAT.—Himalayan Terai, from Central Nepal to the extreme eastern corner of the valley of Assam.

"About three centuries ago this animal existed on the banks of the Indus. The Indian rhinoceros inhabits by preference heavy grass jungle, rarely entering forest. In this respect it differs from its ally Sondaicus, which is a forest-loving species, and even frequents mountainous countries. It is still numerous in the mighty grass jungles which extend along the foot of the Eastern Himalayas from their slopes to the banks of the Brahmaputra. It is yearly becoming more scarce in the Nepal Terai, but is found there from Rohilkund to the Bhootan Doars."

DESCRIPTION.—The accompanying outline sketch, taken from Nature for April 1874, will give a better idea of the animal than a mere verbal description:—

"For convenience of description I will divide the body into five segments—the head, the cervical, the scapular, the abdominal, and the gluteal. At the junction of the head with the neck is a large deep collar or ruff or fold of skin, which gives a very peculiar appearance to the animal. Behind this is a second similar but smaller ruff, which does not hang so low down from the throat as the first. On the dorsal surface it transversely crosses the nape. It is then continued down angularly to about the centre of the anterior edge of the scapular shield, where it forms an obtuse angle with its posterior but major half. It is at the point where it forms this angle that it gives off what I call the cervical fold, which forms the boundary of the top front edge of the scapular shield, but is lost at a point in the shoulder nearly over the centre of the fore limb.

"The scapular shield is a thick cuirass-like plate of skin, studded with round projections about the size of a shilling, and bearing much resemblance to the heads of bolts by which the shield was riveted to the body, and hence called 'boiler-bolt tubercules.' This shield is often removed from the carcase of a slain rhinoceros as a trophy, 'and it is in its centre, but slightly low, that the fatal spot lies which will take him in the heart' (Pollock).

"Between the scapular and the gluteal shields lies the abdominal segment. It calls for no particular description, except that the tubercles here are very much flatter and smaller than on either segments three and four. They are here about the size of a four-anna piece, and they seem to be crowded along the centre line of the body, while the dorsal surface is nearly free from them, and smooth.

"We next come to the gluteal segment. It is in this portion that the boiler-bolt tubercles attain their greatest development, some of them being perhaps three-tenths of an inch high.

"The gluteal segment is laterally crossed by three ridges of skin. The first, which is the only one indicated in the drawing, goes right across the buttock. In some animals there is an indication of a second below this, and about fourteen inches lower down a third, which only goes about a quarter of the way across. The tail is almost concealed in a deep groove, in which lie the perineum, &c. Both the front and hind limb from the point at which they project from the body are finely covered with reticulated skin, forming pentagonal and hexagonal scales, very much as in R. Sondaicus, only much finer and less prominent.

"The Indian rhinoceros has the same habit as the African species of depositing its droppings in one spot till they form huge mounds, which the animal levels with its horns. It is probable that this rhinoceros was found throughout the plains of the N.W. Provinces in unreclaimed spots as late as the fifth or sixth century. According to the observation of Dr. Andrew Smith in South Africa these huge pachyderms do not absolutely require for their support the dense tropical vegetation we should think necessary to supply food to such huge beasts. This gentleman saw over fifty of them in one day in an open country covered with short grass and thorn-bushes about four feet high. From the affinities of the fauna of the N.W. Provinces, which are strongly African, it is probable that the plains of the N.W. Provinces were rather covered with scrubby open jungles and grass than with tropical primeval forests.

"Here and there belts of Dhak (Butea frondosa) were found, and in favoured spots doubtless other tree jungle, but it is improbable that primeval forest has existed since the depression of the Indo-Gangetic plain."—J. Cockburn, MS.

The rhinoceros is supposed to be a very long-lived animal. Dr. Gray ('P. Z. S.' 1867. p. 1011) states on the authority of Mr. Blyth that a pair lived in the Barrackpore Park for forty-five years. They were exactly alike in size and general appearance; they never bred. There is no difference in the horns or form of the skull in the two sexes (Blyth, 'J. A. S. B.' vol. xxxi. p. 155).

NO. 430. RHINOCEROS SONDAICUSThe Javan Rhinoceros (Jerdon's No. 213)

NATIVE NAMES.—The same as last in Hindi; Khyen-hsen, Burmese; Warak, Javanese; Badak, Malayan.

HABITAT.—"The Bengal Sunderbunds, Tipperah, the swamps at the base of the Garo, Khasia, and Naga Hills" (Pollock). "Munipurf, extending into the western provinces of China, southward into Burmah, the Malayan peninsula; Sumatra, Java, and Borneo" (J. Cockburn, MS.).

DESCRIPTION.—"Folds somewhat on the same plan as in Indicus, one marked distinction being that the lateral shoulder fold is continued upward over the back of the neck to form an independent saddle-shaped shield on the nape. The whole body covered with pentagonal or hexagonal warty insulæ. Females hornless" (J. Cockburn, MS.). Males with one horn.

SIZE.—Mr. Cockburn gives the following measurements of a female, which he states is the largest recorded specimen: "Length of body (head and body?), 12 feet 3 inches; tail, 2 feet 4½ inches; height, 5 feet 6 inches." Dr. Jerdon gives: "Length 7 to 8 feet; height, 3½ to 3¾ feet;" and he calls the animal "the lesser Indian rhinoceros," whereas Mr. Cockburn's measurement gives an animal somewhat longer, though not so high as the largest recorded specimen of Indicus. Blyth again writes ('Mammals of Burmah,' see 'J. A. S. B.' vol. xliv. part ii. 1875, p. 50): "It is about a third smaller than R. Indicus, from which it is readily distinguished by having the tubercles of the hide uniformly of the same small size, and also by having a fold or plait of the skin crossing the nape in addition to that behind the shoulder-blades."

This rhinoceros seems to be found at all elevations, like the Sumatran one which was found by General Fytche at an altitude of 4000 feet; it is much more of a forester than the last. Blyth and Jerdon suppose it to be the same as the species hunted by the Moghul Emperor Baber on the banks of the Indus.

GENUS CERATORHINUS

"The skin divided into shields by deep folds; the lumbar fold rudimentary, short, only occupying the middle of the space between the groin and the back; horns two, the front longer, curved backward, the hinder small; conical skull; forehead narrow, flat; the upper part of the nose on each side of the horns narrow, rounded, sub-cylindrical; the occipital region erect, the part near the condyles rather concave; the occipital condyle short, broad, oblong, placed obliquely inferior, scarcely prominent; lachrymal bone very large, irregular shaped."—Dr. Gray, 'P. Z. S.' 1867, p. 1021.

NO. 431. RHINOCEROS vel CERATORHINUS (CROSSI?) LASIOTISThe Ear-fringed Rhinoceros

HABITAT.—Arakan, Tenasserim provinces; one was caught near Chittagong in 1868.

DESCRIPTION.—A thinner hide than with the preceding, and not tuberculated; the folds also are fewer in number; there is one great groove behind the shoulder-blades, and a less conspicuous one on the flank, and some slight folds about the neck and top of the limbs; the horns are two in number, the posterior one being the centre of the nose behind the anterior one, and almost over the anterior corner of the eye; the body (of a young specimen) is covered with long, fine, reddish hair, and the posterior margins of the ears have very long fringes of the same; the tail is short and hairy.

A young specimen of this animal (of which there is an excellent coloured plate in 'P. Z. S.' 1872, p. 494) was captured in 1868 in Chittagong. She had got into a quicksand, and had exhausted herself by floundering about. The natives contrived to attach two ropes to her neck, and, hauling her out, managed to make her fast to a tree. Next morning they found her so refreshed and vigorous that they were afraid to do anything more to her, and so sent messengers to the magistrate of Chittagong to report the capture. The same evening Captain Hood and Mr. Wickes started with eight elephants to secure the prize, and after a march of sixteen hours to the south of Chittagong, they came up to the animal. The elephants at first sight bolted, but were brought back by considerable exertion, and the rhinoceros was made fast to one by a rope. The poor creature roared with fright, and a second stampede ensued, in which luckily the rope slipped off the leg of the rhinoceros to which it was attached. Ultimately she was secured between two elephants and marched into Chittagong, where she soon got very tame. Eventually she was sent to England, and was purchased by the Zoological Society for £1250—a very handsome price, owing doubtless to the rarity of the specimen.

NO. 432. RHINOCEROS vel CERATORHINUS SUMATRENSISThe Sumatran Rhinoceros

NATIVE NAMES.—Kyen-shan, Burmese; Bodok, Malayan.

HABITAT.—Tenasserim provinces; Burmah, extending into Siam; the Malayan peninsula and Sumatra.

DESCRIPTION.—A smaller animal than the preceding, with a hard, black, rough, bristly skin; a deep fold behind the shoulder; ears set closer than in the last species, and filled with black hair internally; the muzzle in front of the first horn is broader; the horns are two in number, and attain a good size, curving, but slightly, backward; the tail is conspicuously longer than in R. lasiotis, and is tapering and not tufted. There is a well drawn and coloured plate of this species in the 'Proceedings of the Zoological Society' for 1872, p. 794, as also several engravings showing the heads of the two animals in juxtaposition.

SIZE.—About 3 feet 8 inches in height at the shoulder.

At first it was considered that R. lasiotis was of this species, and as such it was described and sent to England; but on the subsequent arrival of a genuine R. Sumatrensis from Malacca it was apparent that R. lasiotis was quite distinct. The latter is of larger size, lighter colour, with wide-set ears and a tufted tail. The former is smaller, darker, with narrow-set ears and a long tapering semi-nude tail.31 The Society paid Mr. Jamrach £600 in 1872 for the female specimen from Malacca, which settled the question of separate species. A young R. Sumatrensis was born in the Victoria Docks in London on December 7th, 1872, on board the steamship Orchis. There is a coloured sketch of the little one in the 'P. Z. S.' for 1873, and an interesting account of it and the mother by Mr. Bartlett, the Superintendent of the Society's Gardens. From the circumstances of the capture of the mother it appears that the period of gestation of the rhinoceros is about the same as that of the hippopotamus, viz. seven months.

Although the number of species of living rhinoceros is but few, there are a great many fossil species which show that the animal was more plentiful and in greater variety in prehistoric times.

Remains of the woolly rhinoceros (R. trichorhinus) have been found, like those of the mammoth, imbedded in ice; it was about eleven and a-half feet in length, and its body was covered with woolly hair. A specimen found in 1771 or 1772 was entire, and clothed with skin, but so far decomposed as to prevent more than the head and feet being preserved; remains of other fossil species are found throughout Europe, including Great Britain, and also in India. In 'A Sketch of the History of the Fossil Vertebrata of India' by Mr. R. Lydekker, published in the 'Journal of the Asiatic Society of Bengal,' vol. xlix., 1880, will be found the names of eight species of fossil rhinoceros, inclusive of R. Indicus, which is found in recent alluvia—it is found with two others in the Pleistocene formation, and five others are from the Pleiomiocene.

SUB-ORDER ARTIODACTYLA

We now come to the second division, and a very large one, of the UNGULATA, which in itself is again subdivided into non-ruminants and ruminants. The former comprises the pigs of the Old and the peccaries of the New World and the hippopotami; the latter contains the camels, llamas, deerlets, oxen, antelope, and deer. In the Artiodactyla the toes are even on all feet, being normally four (perfect and rudimentary) with the exception of the camel, giraffe and a few antelope, in which two only are present. To understand the subject thoroughly one must compare the fore-foot of a deer or pig with our own hand; what we call the knee of the former is merely our wrist. The bones which run through the palm of the hand to the knuckles are the metacarpals; they are five in number, corresponding with the thumb and four fingers. In the Artiodactyla—or, I should say, in the Ungulata generally—the thumb is entirely wanting; in the Artiodactyla the fore and little fingers are shorter, rudimentary, or entirely wanting, and the two centre metacarpals, the middle and ring fingers are prolonged into what we call the leg below the knee in these animals, which consist of separate or fused bones terminated by the usual three joints of the finger, on the last of which is placed the hoof.