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A History of North American Birds, Land Birds. Volume 2
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Asyndesmus, Coues, Pr. A. N. S. 1866, 55. (Type, Picus torquatus.)

Gen. Char. Bill about equal to the head; broader than high at the base, but becoming compressed immediately anterior to the commencement of the gonys. Culmen and gonys with a moderately decided angular ridge; both decidedly curved from the very base. A rather prominent acute ridge commences at the base of the mandible, a little below the ridge of the culmen, and proceeds but a short distance anterior to the nostrils (about one third of the way), when it sinks down, and the bill is then smooth. The lateral outlines are gently concave from the basal two thirds; then gently convex to the tip, which does not exhibit any abrupt bevelling. Nostrils open, broadly oval; not concealed by the feathers, nor entirely basal. Fork of chin less than half lower jaw. The outer pair of toes equal. Wings long, broad; lengthened. Tail-feathers broad, with lengthened points.

The species all have the back black, without any spots or streaks anywhere.


Melanerpes erythrocephalus.

883


Dr. Coues places M. torquatus in a new genus, Asyndesmus, characterized by a peculiar texture of the under part and nuchal collar, in which the fibres are disconnected on their terminal portion, enlarged and stiffened, almost bristle-like; otherwise the characters are much as in Melanerpes. It should, however, be noted, that the feathers of the red portion of the head in the other species have the same texture.

Species and Varieties

A. Sexes similar. Young very different from the adult.

M. torquatus. Feathers of the lower parts, as well as of frontal, lateral, and under portions of the head, with the fibres bristle-like. (Asyndesmus, Coues.) Upper parts wholly uniform, continuous, very metallic blackish-green. Adult. Forehead, lores, cheeks, and chin deep crimson, of a burnt-carmine tint; jugulum, breast, and a ring entirely around the nape, grayish-white; abdomen light carmine. Back glossed with purplish-bronze. Young without the red of the head, and lacking the grayish nuchal collar; abdomen only tinged with red, no purple or bronze tints above. Wing, 6.70; tail, 4.50. Hab. Western Province of the United States, from the Black Hills to the Pacific.

M. erythrocephalus. Feathers generally soft, blended; those of the whole head and neck with stiffened and bristle-like fibres in the adult. Secondaries, rump, and upper tail-coverts, with whole lower parts from the neck, continuous pure white. Two lateral tail-feathers tipped with white. Adult. Whole head and neck bright venous-crimson or blood-red, with a black convex posterior border across the jugulum; back, wings, and tail glossy blue-black. Young. Head and neck grayish, streaked with dusky; back and scapulars grayish, spotted with black; secondaries with two or three black bands; breast tinged with grayish, and with sparse dusky streaks. Wing, 5.90; tail, 3.90. Hab. Eastern Province of the United States, west to the Rocky Mountains.

B. Sexes dissimilar; young like the adult.

M. formicivorus. Forehead and a broad crescent across the middle of the throat (the two areas connected by a narrow strip across the lore), white, more or less tinged with sulphur-yellow. Rump, upper tail-coverts, abdomen, sides, and crissum, with patch on base of primaries, pure white, the sides and breast with black streaks. Other portions glossy blue-black.

♂. Whole crown and nape carmine. ♀ with the occiput and nape alone red.

More than the anterior half of the pectoral band immaculate

♀ with the white frontal, black coronal, and red occipital bands of about equal width. Forehead and throat only slightly tinged with sulphur-yellow. Wing, 5.80; tail, 3.90; bill, 1.27. Hab. Pacific Province of United States, and Northern and Western Mexico … var. formicivorus.

♀ with the white frontal band only about half as wide as the black coronal, which is only about half as wide as the red occipital, band or patch. Forehead and throat bright sulphur-yellow. Wing, 5.40; tail, 3.65; bill, 1.23. Hab. Lower California … var. angustifrons.

Nearly the whole of the black pectoral band variegated with white streaks

Relative width of the white, black, and red areas on the crown as in formicivorus. Wing, 5.50; tail, 3.75; bill, 1.22. Hab. Middle America, south of Orizaba and Mirador … var. striatipectus.131

♂. Nape, only, red (as in females of preceding races); ♀ without any red.

Whole breast streaked, the black and white being in about equal amount. Wing, 5.70; tail, 3.90; bill, 1.20. Hab. New Granada … var. flavigula.132

Melanerpes torquatus, BonapLEWIS’S WOODPECKER

Picus torquatus, Wilson, Am. Orn. III, 1811, 31, pl. xx.—Wagler, Syst. Av. 1827, No. 82.—Aud. Orn. Biog. V, 1839, 176, pl. ccccxvi.—Ib. Birds Amer. IV, 1842, 280, pl. cclxxii.—Sundevall, Consp. 51. Melanerpes torquatus, Bp. Consp. 1850, 115.—Heermann, J. A. N. Sc. Phil. 2d ser. II, 1853, 270.—Newberry, Zoöl. Cal. & Or. Route, 90, in P. R. R. Surv. VI, 1857.—Baird, Birds N. Am. 115.—Cooper & Suckley, 161.—Cassin. Pr. A. N. S. 1863, 327.—Lord, Pr. R. A. Inst. IV, 1864, 112 (nesting).—Cooper, Orn. Cal. 1, 1870, 406. Picus montanus, Ord. in Guthrie’s Geog. 2d Am. ed. II, 1815, 316. Picus lewisii, Drapiez. (Gray.) Asyndesmus torquatus, Coues, Pr. A. N. S. 1866, 55.

Sp. Char. Feathers on the under parts bristle-like. Fourth quill longest; then third and fifth. Above dark glossy-green. Breast, lower part of the neck, and a narrow collar all round, hoary grayish-white. Around the base of the bill and sides of the head to behind the eyes, dark crimson. Belly blood-red, streaked finely with hoary whitish. Wings and tail entirely uniform dark glossy-green. Female similar. Length about 10.50; wing, 6.50. Young without the nuchal collar, and the red of head replaced by black.

Hab. Western America from Black Hills to Pacific.

The peculiarities in the feathers of the under parts have already been adverted to. This structure appears to be essentially connected with the red feathers, since these have the same texture in the other species of the genus, wherever the color occurs. The remark may perhaps apply generally to the red feathers of most, if not all, Woodpeckers, and may be connected with some chemical or physical condition yet to be determined.

Habits. Lewis’s Woodpecker would seem to have a distribution throughout the Pacific Coast, from the sea-shore to the mountains, and from Puget Sound to the Gulf of California, and extending to the eastern border of the Great Plains, within the limits of the United States. They were first observed by Messrs. Lewis and Clarke, in their memorable journey to the Pacific. Subsequently Mr. Nuttall met with them in his westward journey, in the central chain of the Rocky Mountains. This was in the month of July. Among the cedar and pine woods of Bear River, on the edge of Upper California, he found them inhabiting the decayed trunks of the pine-trees, and already feeding their young. Afterwards, at the close of August, he met them in flocks on the plains, sixty miles up the Wahlamet. He describes them as very unlike Woodpeckers in their habits, perching in dense flocks, like Starlings, neither climbing branches nor tapping in the manner of their tribe, but darting after insects and devouring berries, like Thrushes. He saw them but seldom, either in the dense forests of the Columbia or in any settled part of California.

Townsend speaks of their arriving about the first of May on Bear River and the Columbia. Both sexes incubate, according to his observations.

Dr. Gambel first observed this Woodpecker in a belt of oak timber near the Mission of St. Gabriel, in California, and states that it was abundant. He also describes its habits as peculiar, and unlike the generality of Woodpeckers. Dr. Heermann, too, speaks of finding it in all the parts of California which he visited. Dr. Newberry, in his Notes on the zoölogy of Lieutenant Williamson’s expedition, refers to it as most unlike the California Woodpecker in the region it occupies and in its retiring habits. He describes it as seeming to choose, for its favorite haunts, the evergreen forests upon the rocky declivities of the Cascade and Rocky Mountains. He first observed it in Northern California, but subsequently noticed it in the mountains all the way to the Columbia. Though often seen in low elevations, it was evidently alpine in its preferences, and was found most frequently near the line of perpetual snow; and when crossing the snow lines, in the mountain-passes, it was often observed flying far above the party. He describes it as being always shy, and difficult to shoot.

Dr. S. W. Woodhouse describes this species as being common in the Indian Territory and in New Mexico; while Dr. Cooper, in his Report on the zoölogy of Washington Territory, speaks of it as being common, during summer, in all the interior districts, but seldom or never approaching the coast. It arrives at Puget Sound early in May, and some even remain, during mild winters, in the Territory. According to his account, it burrows holes for its nests at all heights from the ground, but commonly in dead trees. The eggs are described as pure white, and, when fresh, translucent, like those of all the Woodpecker tribe, and hardly distinguishable in size and general appearance from those of the Golden-winged Woodpecker (Colaptes auratus). Its harsh call is rarely uttered in summer, when it seems to seek concealment for itself and nest. The flocks of young, which in fall associate together to the number of eight or ten, are more noisy. Dr. Suckley, in the same Report (page 162), speaks of this Woodpecker as being very abundant throughout the more open portions of the timbered region of the northwest coast, preferring oak openings and groves. At Fort Dalles, on the Columbia, they are extremely numerous, not only breeding there during summer, but also found as winter residents. Their breeding-places are generally holes in oak and other trees, which, from the appearance of all he examined, seemed to have been excavated for the purpose. At Puget Sound this species was found less frequently than at Fort Dalles, on the Columbia. At the latter place they were constant winter residents. Dr. Suckley also speaks of them as being semi-gregarious in their habits.

Mr. Lord thinks that this Woodpecker is not to be met with west of the Cascade Mountains, but says it is very often found between the Cascades and the Rocky Mountains, where it frequents the open timber. The habits and modes of flight of this bird, he states, are not the least like a Woodpecker’s. It flies with a heavy flapping motion, much like a Jay, feeds a good deal on the ground, and chases insects on the wing like a Shrike or a Kingbird. Whilst mating they assemble in large numbers, and keep up a continual, loud, chattering noise. They arrive at Colville in April, begin nesting in May, and leave again in October. The nest is in a hole in a dead pine-tree, usually at a considerable height from the ground.

Dr. Coues says this bird is very common at Fort Whipple, in Arizona, where it remained in moult until November.

Mr. J. A. Allen found this the most numerous of the Picidæ in Colorado Territory. He also states that it differs considerably in its habits from all the other Woodpeckers. He frequently noticed it rising high into the air almost vertically, and to a great height, apparently in pursuit of insects, and descending again as abruptly, to repeat the same manœuvre. It was met with by Mr. Ridgway in the Sacramento Valley, along the eastern base of the Sierra Nevada, and in the East Humboldt Mountains. In the first-mentioned locality it was the most abundant Woodpecker, and inhabited the scattered oaks of the plains. In the second region it was very abundant—perhaps more so than any other species—among the scattered pines along the very base of the eastern slope; and in the last-mentioned place was observed on a few occasions among the tall aspens bordering the streams in the lower portions of the cañons. In its habits it is described as approaching most closely to our common Red-headed Woodpecker (M. erythrocephalus), but possessing many very distinctive peculiarities. In the character of its notes it quite closely approximates to our common Redhead, but they are weaker and of a more twittering character; and in its lively playful disposition it even exceeds it. It has a very peculiar and characteristic habit of ascending high into the air, and taking a strange, floating flight, seemingly laborious, as if struggling against the wind, and then descending in broad circles to the trees.

The eggs are more spherical than are usually those of the Colaptes auratus, are of a beautiful crystalline whiteness, and measure 1.10 inches in length and .92 of an inch in breadth.

Melanerpes erythrocephalus, SwainsonRED-HEADED WOODPECKER

Picus erythrocephalus, Linn. Syst. Nat. I, 1766, 174.—Vieillot, Ois. Am. Sept. II, 1807, 60, pl. cxii, cxiii.—Wilson, Am. Orn. I, 1810, 142, pl. ix, fig. 1.—Wagler, Syst. Av. 1827, No. 14.—Ib. Isis, 1829, 518 (young).—Aud. Orn. Biog. I, 1832, 141; V, 536, pl. xxvii.—Ib. Birds America, IV, 1842, 274, pl. cclxxi.—Max. Cab. J. VI, 1858, 419. Melanerpes erythrocephalus, Sw. F. B. A. II, 1831, 316.—Bon. List, 1838.—Ib. Conspectus, 1850, 115.—Gambel, J. Ac. Nat. Sc. Ph. 2d ser. I, 1847, 55.—Baird, Birds N. Am. 1858, 113.—Scl. Cat. 1862, 340.—Samuels, 102.—Cooper, Orn. Cal. 1, 1870, 402.—Allen, B. E. Fla. 307. Picus obscurus, Gm. I, 1788, 429 (young).—Red-headed Woodpecker, Pennant, Kalm, Latham. White-rumped Woodpecker, Latham.

Sp. Char. Head and neck all round crimson-red, margined by a narrow crescent of black on the upper part of the breast. Back, primary quills, and tail bluish-black. Under parts generally, a broad band across the middle of the wing, and the rump, white. The female is not different. Length, about 9.75; wing, 5.50. Bill bluish-white, darker terminally; iris chestnut; feet olive-gray. Young without any red, the head and neck being grayish streaked with dusky; breast with an ashy tinge, and streaked sparsely with dusky; secondaries with two or three bands of black; dorsal region clouded with grayish.

Hab. Eastern Province of United States to base of Rocky Mountains, sometimes straggling westward to coast of California (Gambel). Salt Lake City, Utah (Ridgway). Other localities: Nueces to Brazos, Texas (Dresser, Ibis, 1865, 469, breeds).

Western specimens frequently have the abdomen strongly tinged with salmon-red, or orange-red, and are generally more deeply colored than eastern.

Habits. The Red-headed Woodpecker is one of the most familiar birds of this family, and ranges over a wide extent of territory. Excepting where it has been exterminated by the persecutions of indiscriminate destroyers, it is everywhere a very abundant species. Once common, it is now rarely met with in the neighborhood of Boston, though in the western part of Massachusetts it is still to be found. In the collections of the Smithsonian Institution are specimens from Pennsylvania, Wyoming, Nebraska, Kansas, Missouri, Louisiana, the Indian Territory, etc. Sir John Richardson speaks of it as ranging in summer as far north as the northern shores of Lake Huron. He also remarks that in the Hudson Bay Museum there is a specimen from the banks of the Columbia River. Dr. Gambel, in his paper on the birds of California, states that he saw many of them in a belt of oak timber near the Mission of St. Gabriel. As, however, Dr. Heermann did not meet with it in California, and as no other collector has obtained specimens in that State, this is probably a mistake. With the exception of Dr. Woodhouse, who speaks of having found this species in the Indian Territory and in Texas, it is not mentioned by any of the government exploring parties. It may therefore be assigned a range extending, in summer, as far north as Labrador, and westward to the eastern slopes of the Rocky Mountains. Throughout the year it is a permanent resident only of the more southern States, where it is, however, much less abundant in summer than it is in Pennsylvania.

Wilson, at the time of his writing (1808), speaks of finding several of the nests of this Woodpecker within the boundaries of the then city of Philadelphia, two of them being in buttonwood-trees and one in the decayed limb of an elm. The parent birds made regular excursions to the woods beyond the Schuylkill, and preserved a silence and circumspection in visiting their nest entirely unlike their habits in their wilder places of residence. The species is altogether migratory, visiting the Middle and Northern States early in May and leaving in October. It begins the construction of its nest almost immediately after its first appearance, as with other members of its family, by excavations made in the trunk or larger limbs of trees, depositing six white eggs on the bare wood. The cavities for their nests are made almost exclusively in dead wood, rarely, if ever, in the living portion of the tree. In Texas, Louisiana, Kentucky, and the Carolinas, they have two broods in a season, but farther north than this they rarely raise more than one. Their eggs are usually six in number, and, like all the eggs of this family, are pure white and translucent when fresh. They vary a little in their shape, but are usually slightly more oval and less spherical than those of several other species. Mr. Nuttall speaks of the eggs of this bird as being said to be marked at the larger end with reddish spots. I have never met with any thus marked, and as Mr. Nuttall does not give it as from his own observations I have no doubt that it is a mistake. Mr. Paine, of Randolph, Vt., writes that he has only seen a single specimen of this Woodpecker in that part of Vermont, while on the western side of the Green Mountains they are said to be very common. He adds that it is a tradition among his older neighbors that these Woodpeckers were formerly everywhere known throughout all portions of the State.

Mr. Ridgway saw a single individual of this species in the outskirts of Salt Lake City, in July, 1869.

Their eggs vary both in size and in shape, from a spherical to an oblong-oval, the latter being the more usual. Their length varies from 1.10 to 1.15 inches, and their breadth from .80 to .90 of an inch.

Melanerpes formicivorus, var. formicivorus, BonapCALIFORNIA WOODPECKER

Picus formicivorus, Swainson, Birds Mex. in Philos. Mag. I, 1827, 439 (Mexico).—Vigors, Zoöl. Blossom, 1839, 23 (Monterey).—Nuttall, Man. I, 2d ed. 1840. Melanerpes formicivorus, Bp. Conspectus, 1850, 115.—Heermann, J. A. N. Sc. Phil. 2d series, II, 1853, 270.—Cassin, Illust. II, 1853, 11, pl. ii.—Newberry, Zoöl. Cal. & Oregon Route, 90, P. R. R. Surv. VI, 1857.—Baird, Birds N. Am. 1868, 114.—Sumichrast, Mem. Bost. Soc. I, 1865, 562 (correcting an error of Saussure).—Cassin, Pr. A. N. S. 63, 328.—Heermann, P. R. R. X, 58 (nesting).—Baird, Rep. M. Bound. II, Birds, 6.—Sclater, Pr. Z. S. 1858, 305 (Oaxaca).—Ib. Ibis, 137 (Honduras).—Cab. Jour. 1862, 322 (Costa Rica).—Coues, Pr. A. N. S. 1866, 55.—Cooper, Orn. Cal. I, 1870, 403. Picus melanopogon, Temminck, Pl. Color. IV, (1829? pl. ccccli.—Wagler, Isis, 1829, v, 515.—Sundevall, Consp. 51.

Melanerpes formicivorus.


Sp. Char. Fourth quill longest, third a little shorter. Above and on the anterior half of the body, glossy bluish or greenish black; the top of the head and a short occipital crest red. A white patch on the forehead, connecting with a broad crescentic collar on the upper part of the neck by a narrow isthmus, white tinged with sulphur-yellow. Belly, rump, bases of primaries, and inner edges of the outer quills, white. Tail-feathers uniform black. Female with the red confined to the occipital crest, the rest replaced by greenish-black; the three patches white, black, and red, very sharply defined, and about equal. Length about 9.50; wing, 6.00; tail, 3.75.

Hab. Pacific Coast region of the United States and south; in Northern Mexico, eastward almost to the Gulf of Mexico; also on the Upper Rio Grande; south to Costa Rica. Localities: Oaxaca (Scl. P. Z. S. 1858, 305); Cordova (Scl. 1856, 307); Guatemala (Scl. Ibis, I, 137); Honduras (Scl. Cat. 341); Costa Rica (Cab. J. 1862, 322); W. Arizona (Coues, P. A. N. S. 1866, 55).

In most specimens one or two red feathers may be detected in the black of the breast just behind the sulphur-yellow crescent. The white of the breast is streaked with black; the posterior portion of the black of the breast and anterior belly streaked with white. The white of the wing only shows externally as a patch at the base of the primaries.


PLATE LIII.


1. Melanerpes formicivorus. ♂ Cal., 5495.


2. Melanerpes formicivorus. ♀ Cal., 25035.


3. Melanerpes angustifrons. ♂ Cape St. Lucas, 25947.


4. Melanerpes angustifrons. ♀ Cape St. Lucas, 25949.


Dr. Coues calls attention to extraordinary differences in the color of the iris, which varies from white to red, blue, yellow, ochraceous, or brown. A mixture of blue, he thinks, indicates immaturity, and a reddish tinge the full spring coloration.

The male of this species has a white forehead extending a little backwards of the anterior edge of the eye, the rest of the top of head to the nape being red. The female has the white forehead, and a quadrate occipito-nuchal red patch, a black band about as broad as the white one separating the latter from the occipital red. The length of the two anterior bands together is decidedly greater than that of the posterior red. In both sexes the jugulum is entirely and continuously black. Anteriorly (generally with a red spot in its anterior edge) and on the feathers of its posterior border only are these elongated white spots, on each side the shaft, the feathers of the breast being streaked centrally with black. The inner webs of the secondaries have an elongated continuous patch of white along their internal edge, with a very slight, almost inappreciable, border of black; this white only very rarely converted partly or entirely into quadrate spots, and that never on the innermost quills marked with white. Specimens from California are very similar to those from the Rocky Mountains and the Rio Grande Valley, except, perhaps, in being larger, with longer and straighter bill.

In M. flavigula from Bogota, the male has the head marked with the red, black, and white (the red much less in extent, however) of the female M. formicivorus, while the female has no red whatever. All, or nearly all, the feathers of the jugulum have the two white spots, and (as pointed out by Reichenbach) the white of the inner webs of the inner quills is entirely converted into a series of non-confluent quadrate spots. The black streaks on the sides and behind appear to be of greater magnitude, and more uniformly distributed. In both species all the tail-feathers are perfectly black.

A Guatemalan bird, received from Mr. Salvin as M. formicivorus,—and indeed all specimens from Orizaba and Mirador to Costa Rica,—agrees in the main with the northern bird, except that all the black feathers of the jugulum have white spots, as in M. flavigula. The outermost tail-feather of Mr. Salvin’s specimen has two narrow transverse whitish bands, and a spot indicating a third, as well as a light tip. The white markings on the inner quills are more like the northern bird, though on the outermost ones there is the same tendency to form spots as in a few northern specimens (as 6,149 from Los Nogales, &c.). The bill is very different from either in being shorter, broader, much stouter, and the culmen more decurved.

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