A History of North American Birds, Land Birds. Volume 3

Juv. Above plumbeous-brown, tinged with fulvous on head, and more or less washed with the same on the rump; frequently the feathers of the back, rump, scapulars, and wings pass into a reddish tinge at the edge; this color is, however, always prevalent on the head, which is conspicuously streaked with dusky. Tail plumbeous-dusky, darker terminally, with five regular light bars, those toward the base ashy, as they approach the end becoming more ochraceous; these bars are more continuous and regular than in the adult female, and are even conspicuous on the middle feathers. Primaries dusky, passing on edge (terminally) into lighter; spots on the inner webs broader than in the female, and pinkish-ochre; outer webs with less conspicuous corresponding spots of the same. Beneath soft ochraceous; spots as in adult female, but less sharply defined; tibiæ not darker than abdomen.

Hab. Entire continent of North America, south to Venezuela and Ecuador; West India Islands.

Localities: Ecuador (high regions in winter, Scl. P. Z. S. 1858, 451); Cuba (Cab. Jour. II, lxxxiii, Gundlach, Sept. 1865, 225); Tobago (Jard. Ann. Mag. 116); S. Texas (Dresser, Ibis, 1865, 323, breeding?); W. Arizona (Coues, Pr. A. N. S. 1866, 42); Costa Rica (Lawr. IX, 134); Venezuela (Scl. & Salv. 1869, 252).

LIST OF SPECIMENS EXAMINED

National Museum, 42; Boston Society, 11; Philadelphia Academy, 10; Museum Comp. Zoöl., 7; New York Museum, 3; G. N. Lawrence, 2; R. Ridgway, 4. Total, 79.

The plumage of the adult male, which is not as often seen as that of the younger stages and adult female, is represented in the Smithsonian Collection by fifteen specimens, from various parts of North America. Of these, an example from Jamaica exhibits the purest shades of color, though agreeing closely with some specimens from the interior of the United States; the cinereous above being very fine, and of a light bluish cast. The upper tail-coverts are tipped with white; the tail is a quarter of an inch longer than in any North American specimen, one half-inch longer than the average; the wing, however, is about the same.

A specimen from Santa Clara, California (4,475, Dr. J. G. Cooper), like most of those from the Pacific coast, has the cinereous very dark above, while beneath the ochraceous is everywhere prevalent; the flanks are strongly tinged with blue; the black bars of the tail are much broken and irregular. A specimen from Jamaica (24,309, Spanish Town; W. T. March), however, is even darker than this one, the stripes beneath being almost pure black; on the tail black prevails, although the bands are very regular. Nos. 27,061, Fort Good Hope, British America, 43,136, Fort Yukon, Alaska, and 51,305, Mazatlan, Mexico, have the streaks beneath narrow and linear; the ochraceous confined to the tibiæ, which are of a deep shade of this color.

Falco columbarius.

A specimen from Nicaragua (No. 40,957, Chinandega) is like North American examples, but the reddish tinge beneath is scarcely discernible, and confined to the tibiæ, which are but faintly ochraceous; the markings beneath are broad and deep umber, the black shaft-streak distinct.

In the adult female there is as little variation as in the male in plumage, the shade of brown above varying slightly, also the yellowish tinge beneath; the bars on the tail differ in continuity and tint in various specimens, although they are always five in number,—the first concealed by the coverts, the last terminal. In 19,382, Fort Simpson, British America, and 2,706, Yukon, R. Am. (probably very old birds), the light bars are continuous and pale dull ashy.

The young vary about the same as adults. Nos. 19,381, Big Island, Great Slave Lake; 5,483, Petaluma, California; and 3,760, Racine, Wisconsin,—are young males moulting, scattered feathers appearing on the upper parts indicating the future blue plumage.

Var. suckleyi, RidgwayBLACK MERLIN

Sp. Char. A miniature of F. peregrinus, var. pealei. Above, uniform fuliginous-black, the secondaries and tail-feathers very narrowly but sharply tipped with white, and the primaries passing into whitish on their terminal margin; nuchal region with concealed spotting of pale rusty or dingy whitish. Beneath, longitudinally striped with fuliginous-black, or dark sooty-brown, and pale ochraceous; the former predominating on the breast, the latter prevailing on the throat and anal region. Sides and flanks nearly uniform dusky, with roundish white spots on both webs; lower tail-coverts with a broad sagittate spot of dusky on each feather. Lining of the wing fuliginous-dusky, with sparse, small roundish spots of white. Inner webs of primaries plain dusky, without spots, or else with them only faintly indicated. Tail plain dusky-black, narrowly tipped with white, and without any bands, or else with them only faintly indicated.

Male (No. 4,477, Shoalwater Bay, Washington Territory; J. G. Cooper). Wing, 7.35; tail, 5.25; culmen, .50; tarsus, 1.30; middle toe, 1.25.

Female (No. 5,832, Fort Steilacoom, Washington Territory, September, 1856; Dr. George Suckley). Wing, 8.50; tail, 5.70; culmen, .55; tarsus, 1.62; middle toe, 1.35.

Hab. Coast region of Northern California, Oregon, and Washington Territory (probably northward to Alaska). Puget Sound, Steilacoom, Yreka, California (Oct.), and Shoalwater Bay (National Museum).

The plumage of this race is the chief point wherein it differs from the other forms of the species; and in its peculiarities we find just what should be expected from the Oregon region, merely representing as it does the melanistic condition so frequently observable in birds from the northwest coast.

The upper parts are unicolored, being continuous blackish-plumbeous from head to tail. The tail is tipped with white, but the bars are very faintly indicated, being in No. 4,499 altogether wanting, while in 21,333 they can scarcely be discovered, and only four are indicated; in the others there is the usual number, but they are very obsolete. In No. 4,499, the most extreme example, the spots on the inner webs of the primaries are also wanting; the sides of the head are very thickly streaked, the black predominating, leaving the superciliary stripe ill-defined; the throat is streaked, and the other dark markings beneath are so exaggerated that they cover all portions, and give the prevailing color; the under tail-coverts have broad central cordate black spots.

Another specimen from this region (4,476, Puget Sound) is similar, but the spots on primaries are conspicuous, as in examples of the typical style; indeed, except in the most extreme cases, these spots will always be found indicated, leading us to the unavoidable conclusion that the specimens in question represent merely the fuliginous condition of the common species; not the condition of melanism, but the peculiar darkened plumage characteristic of many birds of the northwest coast, the habitat of the present bird; it should then be considered as rather a geographical race, co-equal to the Falco gyrfalco, var. labradora, F. peregrinus, var. pealei, and other forms, and not confounded with the individual condition of melanism, as seen in certain species of Buteones.

LIST OF SPECIMENS EXAMINED

National Museum, 6.

Second quill longest; first quill equal to, a little shorter than, or a little longer than, the fourth.

Var. richardsoni, RidgwayRICHARDSON’S MERLIN

Falco æsalon, Rich. & Swains. F. B. A. II, pl. xxv, 1831.—Nutt. Man. Orn. II, 558.—Coues, P. A. N. S. Philad. 1866, p. 42 (in text). Falco (Hypotriorchis) richardsoni, Ridgway, P. A. N. S. Philad. Dec. 1870, 145. Falco richardsoni, Coues, Key, 1872, p. 214.

Sp. Char. Adult male like the female and young? The known stages of plumage more like the adult female and young of var. lithofalco (F. æsalon, Auct.) than like var. columbarius.

Adult male (Smithsonian, No. 5,171, mouth of the Vermilion River, near the Missouri, October 25, 1856; Lieutenant Warren, Dr. Hayden). Upper plumage dull earth-brown, each feather grayish-umber centrally, and with a conspicuous black shaft-line. Head above approaching ashy-white anteriorly, the black shaft-streaks being very conspicuous. Secondaries, primary coverts, and primaries margined terminally with dull white; the primary coverts with two transverse series of pale ochraceous spots; outer webs of primaries with spots of the same, corresponding with those on the inner webs. Upper tail-coverts tipped, and spotted beneath the surface, with white. Tail clear drab, much lighter than the primaries, but growing darker terminally, having basally a slightly ashy cast; crossed with six sharply defined, perfectly continuous bands (the last terminal) of ashy-white. Head, frontally, laterally, and beneath,—a collar around the nape (interrupting the brown above),—and the entire lower parts, white, somewhat ochraceous, this most perceptible on the tibiæ; cheeks and ear-coverts with sparse, fine hair-like streaks of black; nuchal collar, jugulum, breast, abdomen, sides, and flanks with a medial linear stripe of clear ochre-brown on each feather; these stripes broadest on the flanks; each stripe with a conspicuously black shaft-streak; tibiæ and lower tail-coverts with fine shaft-streaks of brown, like the broader stripes of the other portions. Chin and throat, only, immaculate. Lining of the wing spotted with ochraceous-white and brown, in about equal amount, the former in spots approaching the shaft. Inner webs of primaries with transverse broad bars of pale ochraceous,—eight on the longest. Wing-formula, 2, 3–4, 1. Wing, 7.70; tail, 5.00; culmen, .50; tarsus, 1.30; middle toe, 1.25; outer, .85; inner, .70; posterior, .50.

Adult female (58,983, Berthoud’s Pass, Rocky Mountains, Colorado Territory; Dr. F. V. Hayden, James Stevenson). Differing in coloration from the male only in the points of detail. Ground-color of the upper parts clear grayish-drab, the feathers with conspicuously black shafts; all the feathers with pairs of rather indistinct rounded ochraceous spots, these most conspicuous on the wings and scapulars. Secondaries crossed with three bands of deeper, more reddish ochraceous. Bands of the tail pure white. In other respects exactly as in the male. Wing-formula, 3, 2–4–1. Wing, 9.00; tail, 6.10; culmen, .55; tarsus, 1.40; middle toe, 1.51.

Young male (40,516, Fort Rice, Dacotah, July 20, 1865; Brig.-Gen. Alfred Sully, U. S. A., S. M. Rothammer). Differing from the adult only in minute details. Upper surface with the rusty borders of the feathers more washed over the general surface; the rusty-ochraceous forms the ground-color of the head,—paler anteriorly, where the black shaft-streaks are very conspicuous; spots on the primary coverts and primaries deep reddish-ochraceous; tail-bands broader than in the adult, and more reddish; the terminal one twice as broad as the rest (.40 of an inch), and almost cream-color in tint. Beneath pale ochraceous, this deepest on the breast and sides; markings as in the adult, but anal region and lower tail-coverts immaculate; the shaft-streaks on the tibiæ, also, scarcely discernible. Wing, 7.00; tail, 4.60.

Hab. Interior regions of North America, between the Mississippi Valley and the Rocky Mountains, from Texas to the Arctic regions.

LIST OF SPECIMENS EXAMINED

National Museum, 10; Museum Comp. Zoöl., 2; R. Ridgway, 3. Total, 15.

Since originally describing this bird, I have seen additional examples, and still consider it as an easily recognized race, not at all difficult to distinguish from columbarius. Now, however, I incline strongly to the theory that it represents merely the light form of the central prairie regions, of the common species; since its characters seem to be so analogous to those of the races of Buteo borealis and Bubo virginianus of the same country. It is doubtful whether some very light-colored adult males, supposed to belong to columbarius, as restricted, should not in reality be referred to this race, as the adult plumage of the male. But having seen no adult males from the region inhabited by the present bird obtained in the breeding-season, I am still in doubt whether the present form ever assumes the blue plumage.

As regards the climatic or regional modifications experienced by the Falco lithofalco on the American continent, the following summary of facts expresses my present views upon the subject. First: examples identical in all respects, or at least presenting no variations beyond those of an individual character, may be found from very widely separated localities; but the theory of explanation is, that individuals of one race may become scattered during their migrations, or wander off from their breeding-places. Second: the Atlantic region, the region of the plains, and the region of the northwest coast, have each a peculiar race, characterized by features which are also distinctive of races of other birds of the same region, namely, very dark—the dark tints intensified, and their area extended—in the northwest coast region; very light—the light markings extended and multiplied—in the middle region; and intermediate in the Atlantic region.

Habits. The distribution of the well-known Pigeon Hawk is very nearly coextensive with the whole of North America. It is found in the breeding-season as far to the north as Fort Anderson, on the Anderson and McKenzie rivers, ranging even to the Arctic coast. Specimens were taken by Mr. Ross at Lapierre House and at Fort Good Hope. Several specimens were taken by Mr. Dall at Nulato, where, he states, it is found all the year round. They were also taken by Bischoff at Kodiak. During the breeding-season it is found as far south as Nova Scotia, New Brunswick, and the northern portions of Maine, and probably Vermont and New York. It is abundant on the Pacific coast.

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1

The whole of the systematic portion of the article on the Raptores, has been prepared by Mr. Ridgway; the biographies, however, are furnished by Dr. Brewer, as usual. The outlines of the skulls and sterna of the Strigidæ, the skulls, sterna, and heads of the Cathartidæ, and the generic outlines of the Falconidæ and Cathartidæ, were drawn by Mr. Ridgway.—S. F. Baird.

2

By Thomas H. Huxley, F. R. S., V. P. Z. S.; Proceedings of the Zoölogical Society of London, 1867, pp. 415–473.

3

By Charles Ludwig Nitzsch. English edition, translated from the German by Dr. Philip Lutley Sclater, and published by the Ray Society of London, 1867.

4

By William McGillivray, A. M.; London, 1840.

5

See Jardine’s Contributions to Ornithology, London, 1849, p. 68; 1850, p. 51; 1851, p. 119; 1852, p. 103; and Transactions of the Zoölogical Society of London, 1862, p. 201.

6

Hand List of Genera and Species of Birds, distinguishing those contained in the British Museum. By George Robert Gray, F. R. S., etc. Part I. Accipitres, Fisserostres, Tenuirostres, and Dentirostres. London, 1869.

7

I have, however, examined the sterna only of Nyctea, Bubo, Otus, Brachyotus, Syrnium, Nyctale, and Glaucidium.

8

My unpublished determinations of the North American species were furnished, by request, to Dr. Coues, for introduction into his “Key of North American Birds”; consequently the names used in these pages are essentially the same as those there employed.

9

This case of the restriction of the American representative of a European or Western Palæarctic species to the western half of the continent has parallel instances among other birds. The American form of Falcolanarius (var. polyagrus), of Corvus corax (var. carnivorus), Pica caudata (var. hudsonica and var. nuttalli) and of Ægialitis cantianus (var. nivosus), are either entirely restricted to the western portion, or else are much more abundant there than in the east. The European genera Cinclus, Coccothraustes, Nucifraga, and Columba have representatives only in the western portion of North America.

Instances of a similar relation between the plants of the Western Province of North America and those of Europe, and more striking likeness between the flora of the Eastern Region and that of Eastern Asia, are beautifully explained in Professor Gray’s interesting and instructive paper entitled “Sequoia, and its History,” an address delivered at the meeting of the American Association for the Advancement of Science, at Dubuque, Iowa, August, 1872. The poverty in the species of tortoises, and richness in lizards, and the peculiarities of the ichthyological fauna, as well as absence of forms of Western North America and Europe, compared with Eastern North America and Eastern Asia, afford other examples of parallelism in other classes of the Animal Kingdom.

10

See Baird, Am. Journ. Arts and Sciences, Vol. XLI, Jan. and March, 1866; Allen, Bull. Mus. Comp. Zoöl. Cambridge, Vol. II, No. 3; and Ridgway, Am. Journ. Arts and Sciences, Vols. IV and V, Dec., 1872, and Jan., 1873.

11

For diagnoses of these geographical races of Strix flammea, see pp. 1339 and 1340.

12

See Allen, Bull. Mus. Comp. Zoöl., Cambridge, Vol. II, No. 3, pp. 338, 339, where these plumages are discussed at length.

13

Strix flammea, var. flammea. Strix flammea, Linn. S. N. I, 133, 1766, et Auct.-Strickl. Orn. Syn. I, 1855, 178. Strix alba, Scop. Ann. 1st, p. 21, 1768.—Gmel. S. N. 293.—Lath.—Daud.—Lep. and Shaw. Strix guttata, Brehm, Vög. Deutschl. p. 106, 1831. Hab. Europe and Africa.

14

Strix flammea, var. guatemalæ, Ridgway. Central American specimens differ very appreciably from Mexican and North American examples, in being considerably darker-colored in the extreme phases of plumage. Eight of eleven specimens convey an impression of decided difference in this respect at a mere casual glance. The extremes of plumage in this series are as follows:—Darkest (No. 40,961, Chimandega, Nicaragua; F. Hicks): The dusky mottling of the upper parts is altogether darker than in any example from Mexico or northward, and prevails, with great uniformity, over the entire surface; the white specks are linear, instead of roundish. On the primaries and tail, the blackish and ochraceous are about equal in extent, the latter color forming five bands on the quills, and four on the tail. The facial circle is bright dark orange above the ears, and the portion below the ears continuous black; the face is reddish-white, strongly tinged with wine-brown, while the spot in front of the eye is deep black. The whole lower parts are deep orange-ochraceous, with numerous irregular specks of dusky, which posteriorly become broken or confused into ragged zigzag transverse mottlings, while on the lower tail-coverts they form irregular transverse bars. Wing-formula, 2, 1–3. Wing, 13.00; tail, 5.90; culmen, .85; tarsus, 2.90; middle toe, 1.50. Lightest (No. 41,252, ♂, San José, Costa Rica, Aug. 23, 1865; José C. Zeledon): The dark tint above, though prevalent, is a continuous wash of grayish, instead of a fine mottling of blackish and white; the white specks are nearly obsolete. The wings are superficially plain grayish, this overlaying the ground-color of ochraceous-orange; and have visible spots only on the primaries, near their shafts. The tail has four rather distinct grayish bands. The facial circle is ochraceous, somewhat darker across the foreneck; the face white, with the ante-orbital spot claret-brown. Entire lower parts immaculate pure white. Wing-formula, 2, 1=3. Wing, 12.30; tail, 5.30; culmen, .70; tarsus, 2.75; middle toe, 1.45.

No. 24,283, Nicaragua, (Captain J. M. Dow,) is like the specimen just described, in the uniform dark wash of the upper parts, but this is deeper; the lower parts, however, are quite different, being ochraceous-orange, instead of pure white.

The remaining five specimens (from San Salvador, Costa Rica, and Nicaragua) are alike, and differ from northern birds in the deeper dark mottling of the upper parts; the white specks very conspicuous, and usually sagittate. The facial circle deep black where it crosses the foreneck. The lower parts vary in color from nearly pure white to deep orange-rufous; the dark markings of the lower surface are larger, more angular, and more transverse than in true pratincola. The wing measures 11.30–13.00; tarsus, 2.55–2.95. The northern form varies from 12.50–13.00 (wing) and 2.50–2.85 (tarsus). It is thus seen that while these southern birds average smaller in general dimensions, they have actually larger feet, the average length of the tarsus being 2.80 in the Central American series, and only 2.60 in the northern series. This exactly coincides with the case of Sturnella, the S. magna var. mexicana of the same region being smaller bodied and shorter winged than var. magna of the United States, but with much larger feet, see p.

15

Strix flammea, var. perlata. Strix perlata, Licht. Verz. Doubl. 59, 1823.—Tschudi, Av. Consp. Wiegm. Archiv. 267, 1844.—Hartt. Syst. Ind. Azara. p. 3.—Max. Beitr. III, 263 (excl. syn.).—Strickl. Orn. Syn. I, 1855, 180 (excl. syn.). Strix flammea, Darwin, Zool. Beag. 34.—Schomb. Verz. Faun. Brit. Guian. p. 732.—Spix, Av. Bras. I, 21.

This is a still further differentiated or more appreciably modified race. It differs in smaller size (wing, 11.70–12.50; tail, 4.80–5.20; tarsus, 2.40–2.75) and more square tail, while the colors also present constant differences. The tail is much lighter compared with the wings, the bands narrower and more sharply defined, though the same in number.

16

Strix flammea, var. furcata. Strix furcata, Temm. Pl. Col. 432, 1838.—D’Orb. Hist. Nat. Cuba Ois. p. 34. Hab. West Indies (Cuba and Jamaica).

This form is the most distinctly characterized of all the races of S. flammea which we have examined. It has the general plumage decidedly lighter and less rufous, while the secondaries and tail are abruptly lighter than the adjacent parts, and usually free from bands, though there are sometimes traces of them.

All the American races of Strix flammea differ very decidedly from the European form (var. flammea) in much larger size. The differences in color are not so appreciable, and there is hardly any certain difference in this respect. The extreme phases, however, appear to be darker in the var. flammea than in the var. pratincola. The supposed differences in the character of the feathers fringing the operculum, insisted on by MacGillivray (History of British Birds, III, 1840, p. 473), I am unable to appreciate, for I cannot find that they differ in the least in the two races. That excellent ornithologist states that in the American “species” the feathers of the operculum are reduced to a simple tube, having neither filaments nor shaft, while in the European bird they are perfect feathers, with all their parts complete. Though this may have been the case with the one or more specimens of pratincola examined by Mr. MacGillivray, I have yet to see an American specimen which has not the feathers of the operculum just as perfectly developed as in European examples.