A History of North American Birds, Land Birds. Volume 1

Highly scansorial; tail of twelve rigid acuminate feathers, whereof the outer pair are short and spurious, concealed between bases of next two pairs. Bill stout, straight, with the tip truncate or acute, not decurved,—an efficient chisel for hammering and boring wood. Tongue vermiform, extensile,19 and barbed. Salivary glands large; hyoidean apparatus peculiar. Nasal tufts usually present. Arboreal. (Woodpeckers.) … Picidæ.

Feet neither syndactyle nor zygodactyle.

Feet semipalmate, of normal ratio of phalanges. Anterior toes connected at base by movable webbing. Hind toe very small, elevated, semilateral. Middle toe produced, its large claw pectinate. Bill fissirostral, with very small, triangular, depressed horny part and immense rictus, reaching below the eyes, furnished with bristles. Rather large. Plumage soft and lax, much variegated … Caprimulgidæ.

Feet scarcely or not semipalmate, of frequently abnormal ratio of phalanges (middle or outer toe, or both, with fewer joints than usual among birds). Hallux very small, elevated, frequently lateral or versatile. Middle toe not produced nor its claw pectinate. Bill much as in the last, but rictus unbristled. Small. Plumage compact, of few simple subdued colors … Cypselidæ.

Family of PSITTACI

To characters of Psittaci add: Cere feathered, concealing the nostrils. Feet granular, rugose. Wings pointed. Tail cuneate. Plumage coarse and dry. Head feathered. Colors green, with yellow and blue … Psittacidæ.20

Analysis of Families of RAPTORES

Feet highly raptorial, with large, strong, sharp, curved, contractile claws, adapted for grasping. Hallux perfectly incumbent, lengthened (more than half as long as the fourth toe), with large claw. Front toes with slight basal webbing between outer or middle ones, or none; outer toe often reversible. Nostrils imperforate. Bill short, stout, not notably contracted in its continuity, with strongly hooked tip; tomia often once-twice toothed or lobed. Head feathered wholly or in greatest part. Lower larynx developed with one pair of muscles. Plumage with or without after-shafts. Cœca present, as a rule, if not always.

Physiognomy peculiar by reason of great lateral expansion and lengthwise shortening of the cranium, causing the eyes to be directed forward. Eyes surrounded by a disc of radiating bristly feathers, in front closely appressed to and hiding the base of the bill, elsewhere bounded by a rim of differently formed feathers. Tomia never toothed or lobed. Nostrils usually at the edge of the cere. Outer toe completely versatile, shorter than the inner toe. Basal phalanx of middle toe not longer than the second, and much shorter than the next. Legs commonly feathered or bristly to or on the toes. Plumage peculiarly soft and lax, without after-shafts; flight perfectly noiseless. Cranial walls widely separated by intervention of spongy diploë. Sternum commonly doubly notched. Chiefly nocturnal … Strigidæ.

Physiognomy not peculiar in any lateral expansion of the cranium; the eyes lateral in direction. No complete facial disc; base of bill not hidden by appressed bristles. Nostrils wholly in the cere. Outer toe rarely versatile, except Pandion, etc.; not shorter than the inner. Basal phalanx of middle toe longer than the second. Legs commonly naked and scutellate or reticulate in some portion of their length; toes always bare and scaly. Plumage compact, usually with after-shafts; flight audible. Cranial walls with little diploë. Sternum commonly single-notched or fenestrate, sometimes entire. Diurnal … Falconidæ.

Feet scarcely raptorial, with lengthened, little curved or contractile, weak, short claws. Hallux elevated, shortened, not more than half as long as the fourth toe, with small claw. Front toes all webbed at base; middle toe lengthened; outer not reversible. Basal phalanx of middle toe longer than either of the succeeding. Nostrils perforate. Bill lengthened and comparatively weak, little hooked, contracted in its continuity; tomia not toothed or lobed. Head naked of feathers in greatest part; sparsely bristly. No lower larnyx developed. No cœca. After-shafts absent … Cathartidæ.

Family of COLUMBÆ

With characters essentially as in Columbæ (exclusive of those peculiar to Diduncudidæ and Dididæ). Plumage without after-shafts; the feathers with thickened, spongy rhachis loosely inserted in the skin. Head small, completely feathered, excepting sometimes a circumorbital space. Tarsi naked or only feathered a little way above. Tail of twelve feathers, or lengthened, cuneate, and of fourteen. (Hallux not perfectly incumbent in Starnænas.) … Columbidæ.

Analysis of Families of GALLINÆ

Hind toe lengthened, insistent. Tail-feathers twelve. Sides of head and throat with naked spaces. Color greenish … Cracidæ.

Hind toe shortened, elevated. Tail-feathers usually fourteen or more. No green.

Large. Tarsi, toes, and nasal fossæ naked. Head bare of feathers, sparsely bristly, with wattles and caruncles. A pectoral tuft of bristly feathers. Tarsi usually spurred in the male. Plumage iridescent … Meleagrididæ.

Medium. Tarsi wholly or in great part, sometimes also the toes, and always the nasal fossæ, feathered. Head completely feathered, excepting a definite papillate strip over the eye. Tail-feathers sixteen or more. Sides of neck usually with lengthened feathers, or a naked distensible area, or both. No spurs. Plumage without iridescence … Tetraonidæ.

Small. Tarsi, toes, and nasal fossæ naked. Head completely feathered. No peculiar feathers or tympanum on sides of neck. No spurs. Plumage not iridescent … Perdicidæ.

Analysis of Families of LIMICOLÆ

Toes not lobate. Tarsi not notably compressed.

Legs extremely long; the tarsus equalling or exceeding the tail, and feet either four-toed and palmate (Recurvirostra), or three-toed and semipalmate (Himantopus); with the bill much longer than the head, very slender, acute, and curved upward … Recurvirostridæ.

Legs moderate, stout. Tarsus shorter than tail. Bill hard, more or less contracted at base, with short nasal fossa, gonydeal angle, and ascending gonys, the tip either compressed and truncate or depressed and acute. Feet three-toed and with basal webbing (Hæmatopus), or four-toed and cleft (Strepsilas) … Hæmatopodidæ.

Legs moderate. Tarsus shorter than tail, reticulate. Hind toe wanting (except in Squatarola, where very small, and in Aphriza). Bill short, straight,—not exceeding the head (generally shorter),—shaped like a pigeon’s, with short, broad, soft nasal fossæ separated by a constriction from the enlarged, obtuse, horny terminal part. Head large, globose, contracting suddenly to the bill. Neck short … Charadriidæ.

Legs moderate. Tarsus shorter than tail, scutellate. Hind toe present. Bill long,—equalling, or oftener exceeding, frequently several times longer than, the head; softish and membranous to the very tip, without constriction in its continuity; straight or variously curved … Scolopacidæ.

Toes lobate. Tarsi notably compressed.

General characters of Scolopacidæ. Body depressed; the under plumage thickened, duck-like. Habits natatorial … Phalaropodidæ.

Analysis of Families of HERODIONES

Hallux lengthened, perfectly incumbent, with large claw. Tarsi scutellate. Middle claw pectinate. Bill perfectly straight, tapering, acute. Loral region definitely naked, continuous with covering of the bill. Head narrow, elongate, tapering … Ardeidæ.

Hallux somewhat reduced, less perfectly incumbent. Tarsi commonly reticulate. Middle claw not pectinate. Lores, gular space and usually more of the head, naked. Bill variously curved or with expanded tip. (Genera Tantalus, Ibis, Mycteria, and Platalea.) … Tantalidæ.

Analysis of Families of ALECTORIDES

Of great stature, with extremely long neck and legs. Part or all of the head bare. Toes much shorter than the tarsi; with basal webbing, but without lobation; hallux very short, highly elevated. Bill equalling or exceeding the head, compressed, perfectly straight, contracted about the middle, with enlarged acute terminal portion; nasal fossæ wide and deep, with large perforate nostrils … Gruidæ.

Size moderate and small; neck and legs comparatively short. Head completely feathered, excepting, in the Coots and Gallinules, a broad horny frontal plate. Toes equalling or exceeding the tarsi, simple or lobate. Bill not constricted in the middle, rather shorter than the head, straight and quite stout; or much longer, regularly slender and decurved, with long nasal fossæ. Nostrils incompletely or not perforate … Rallidæ.

Analysis of Families of LAMELLIROSTRES

Of great stature, with extraordinarily lengthened neck and legs. Bill of unique shape, bent abruptly down from the middle. Tibiæ naked below. Hind toe minute or absent. Wings rounded. Red the chief color … Phœnicopteridæ.

Of moderate size; the neck short, or, when lengthened, not accompanied by co-ordinately lengthened legs, these being always shorter than the wing. Bill straight. Tibiæ feathered below. Hind toe present; well developed and functional, though short … Anatidæ.

[Pg xxviii] Analysis of Families of STEGANOPODES

Bill rather longer than head, cleft to eyes, very stout at base, tapering to the decurved, but not hooked, tip. Nostrils abortive. Gular sac moderate, naked. Wings rather long, pointed. Tail long, stiff, cuneate, twelve to fourteen feathered. Feet nearly beneath centre of equilibrium. General configuration goose-like … Sulidæ.

Bill several times longer than head, slender but strong, depressed, perfectly straight, with small distinct hooked nail at end. Nostrils very small. Gular sac enormous. Mandibular rami meeting only at tip. Wings extremely long, with upward of forty remiges. Tail short, rounded, of twenty or more feathers. Legs beneath centre of equilibrium, extremely short and stout … Pelicanidæ.

Bill about as long as head, stout, straight, scarcely tapering, strongly hooked. Nostrils abortive. Gular sac moderate, but evident; mostly naked. Wings short. Tail large, fan-shaped, scansorial, of twelve to fourteen broad stiff feathers, exposed to the base. Legs inserted far behind centre of equilibrium … Graculidæ.

Bill rather longer than head, slender, perfectly straight, tapering to an acute tip. Gular sac small. Nostrils minute. Wings and tail, and general configuration, as in the last … Plotidæ.

Bill much longer than head, straight, stout, strongly hooked. Nostrils very small. Gular sac well developed. Wings exceedingly long, strong, and pointed. Tail exceedingly long, deeply forked. Feet extraordinarily short; tarsi partly feathered … Tachypetidæ.

Bill about as long as head, straight, stout, tapering to an acute tip. Nostrils small. Gular sac rudimentary, feathered. Wings moderate, pointed. Tail short, but with two central feathers extraordinarily prolonged and filamentous. Feet small, beneath centre of equilibrium … Phæthontidæ.

Analysis of Families of LONGIPENNES

Nostrils not tubular, lateral, perforate. Bill with continuous covering, or only broken by a sort of cere, hooked or straight to the end. Hallux small and elevated, but always present … Laridæ.

Nostrils tubular, disjoined and lateral, or oftenest superior and united in one double-barrelled tube. Covering of bill in several pieces; bill always hooked. Hallux minute, rudimentary, or absent … Procellariidæ.

Analysis of Families of PYGOPODES

Feet four-toed, palmate. Hallux lobate, connected at base with base of inner toe. Tail perfect. Head closely and completely feathered. Nostrils with a depending lobe or flap. Bill straight, compressed, acute … Colymbidæ.

Feet four-toed, lobate. Hallux lobate, free. Tail rudimentary. Head with a naked loral strip and bristly or variously lengthened feathers. Nostrils simple. Bill straight or decurved at end, compressed, acute … Podicipidæ.

Feet three-toed, palmate. Hallux absent. Tail perfect. Head closely feathered or variously crested. Nostrils simple. Bill of indeterminate shape … Alcidæ.

NORTH AMERICAN BIRDS

Family TURDIDÆ.—The Thrushes

The Turdidæ, with the Saxicolidæ and Cinclidæ, form a group closely related, by common characters, and appreciably different from the other Oscines with slender bills and specially insectivorous habits, having, like them, ten primaries (the first much shorter than the second, but nearly always appreciable), and the nostrils uncovered. The great family of Sylvicolidæ, with similar characters of the bill, never present more than nine primaries. The most striking of these common characters is seen in the deeply cleft toes, of which the outer is united by the basal joint alone to the middle toe, while the inner is separated almost to the very base of its first joint.21 The frontal feathers extend, with rare exceptions, to the very nostrils. The bill is elongated and subulate, moderately slender, and usually notched at tip; the culmen moderately curved from the base, and the mouth well provided with bristles, except in a few cases. Usually the scutellæ covering the front and sides of the tarsus are fused into one continuous plate, or else scarcely appreciable, except on the inner edge only; in the Mocking Thrushes they are, however, distinctly marked. The lateral toes are nearly equal, the outer rather the longer. With these as some of the principal characteristics, they may be distinguished from each other as follows:—

Note.—In the present work the length of the tail is measured from the coccyx, inside of the skin, and not, as usually the case, from the base of the quills at their insertion. The wings are measured from the carpal joint, with dividers.

A. Nostrils oval. Loral and frontal feathers with bristly points, or interspersed with bristles; rictus with longer or shorter bristles.

Saxicolidæ. Wings very long and much pointed, reaching beyond the middle of the short square or emarginated tail, and one and a half times or more the length of the latter. The spurious primary very short, the second quill longer than the fourth. In the closed wing the outer secondary reaches only about two thirds the length of longest primary.

Turdidæ. Wings moderate, more rounded, not reaching beyond middle of the often rounded tail, and not more than one and a third the latter, usually more nearly equal. Spurious primary sometimes half the length of second quill; the second quill shorter than the fourth. In the closed wing the outer secondary reaches three fourths or more the length of longest primary.

B. Nostrils linear, in lower edge of nasal membrane. Loral and frontal feathers soft and downy, and no bristles or bristly points whatever about the mouth.

Cinclidæ. Body very short and broad. Wings short, rounded, and concave.

The American Sylviidæ are in some respects very closely related to the Saxicolidæ, but may be distinguished by their much smaller size, more slender and depressed bill, more strongly bristled rictus, etc.; on which account they are more strictly “fly-catchers,” taking their prey in great part on the wing.

Of the three families, the Turdidæ contain a great variety of forms, and exhibit widely different characters, rendering it exceedingly difficult to arrange them in any systematic or regular sequence, or to accurately define their boundaries. In the Birds of North America, the Mocking Thrushes were placed among the Wrens, on account of the distinct tarsal scutellæ, and other characters. We are now, however, inclined to believe, with Dr. Sclater, that their place is with the recognized Turdidæ; and, among other reasons, on the ground of their more deeply cleft toes, and greater extension forward of frontal feathers. The following synopsis of the North American forms will serve the purpose of determining the genera, even if these are not arranged or combined in a strictly natural manner.

A. Turdinæ.—Tarsus covered anteriorly with a continuous plate without scales.

Wings decidedly longer than the tail, which is nearly even. Bill considerably shorter than the head.

First quill usually not one fourth the second. Wings pointed. Tarsus hardly the length of head, but yet longer than middle toe; outstretched toes falling short of tip of tail … Turdus.

B. Miminæ.—Tarsi scutellate anteriorly; scales seven.

Wings decidedly longer than the tail, which is nearly even. Tarsus as long as the head.

Bill decidedly shorter than the head, scarcely notched; wings pointed; first quill less than half the second, third and fourth longest. Claws not peculiar. Bristles prominent. Tarsus considerably longer than middle toe and claw … Oreoscoptes.

Wings decidedly shorter than the tail, which is considerably graduated; first quill half or more than half the second.

Bill notched at tip, shorter than head; straight.

Scutellæ very distinct; gonys straight, or even declining at tip … Mimus.

Scutellæ more or less obsolete; gonys convex, ascending at tip … Galeoscoptes.

Bill not notched at tip, lengthened; sometimes much decurved … Harporhynchus.

Note.—In the Review of American Birds, I., May, 1866, 409, I have advanced the suggestion that the N. American genus Myiadestes, usually placed under the Ampelidæ, really belongs under Turdidæ in a group Myiadestinæ. The relationships are certainly very close, as is shown by the characters given below.

Common Characters.—Tarsi without regular transverse scutellæ, except at lower end. Wings acute, pointed, as long as or longer than tail, which is but slightly graduated. First primary rarely half second, which exceeds the secondaries. Base of quills buffy yellow, as are inner edges. Tail spotted or varied at the end. Young birds with many light spots. Very melodious singers.

Myiadestinæ. Bill short, much depressed; mouth deeply cleft; width at base about equal to the distance from nostril to tip, or greater; commissure more than twice distance from nostrils to tip of bill, and nearly two and a half times length of gonys. Legs weak; tarsi rather longer than middle toe and claw. Tail feathers tapering slightly from base to near tip, giving a slightly cuneate appearance to the tail.

Turdinæ. Bill stouter, more lengthened; narrow at base and more compressed; width at base less than distance from nostril to tip; commissure not more than twice distance from nostrils to tip of bill, and about twice length of gonys. Tarsi stouter, longer than middle toe and claw. Tail feathers widening slightly from base to near tip, giving a parallel-sided or slightly fan-shaped appearance to the tail.

The Miminæ differ, as already mentioned, in the scutellate tarsi: more rounded wings, etc.—S. F. B.

Subfamily TURDINÆ

There are several American genera of Turdinæ not found north of Mexico as yet, although it is not impossible that one of these (Catharus) may hereafter be detected within the limits of the United States. The species of Catharus resemble the North American wood-thrushes (Hylocichla); but the spurious or first primary quill is longer (from one half to one third the second quill), the wings are rounded, not pointed, the tarsus is longer than the head, and the outstretched toes extend beyond the tail. The species to be looked for are C. melpomene and occidentalis.22

The North American species of Turdinæ, while retained under the single genus Turdus, yet constitute several distinct groups, which we may call subgenera.

Genus TURDUS, Linnæus

Turdus, Linnæus, Syst. Nat., ed. 10, 1758, 168. (Type, Turdus viscivorus of Europe.)—Baird, Rev. Am. Birds.

Gen. Char. Bill conical, subulate, shorter than the head; the tip gently decurved and notched (except in Hesperocichla); the rictus with moderate bristles; the wings rather long and pointed, with small first primary (less than one fourth the second); wings considerably longer than the tail, which is firm, nearly even, with broad feathers. Tarsi variable, seldom as long as the skull, the scutellæ fused into a continuous plate, only in rare individual instances showing indications of the lines of separation.

The genus Turdus is very cosmopolitan, occurring nearly throughout the globe, excepting in Australia, and embraces species of highest perfection as singers. In the large number of species known there are many variations in external form, but the transition from one to the other is so gradual as to render it very difficult to separate them into different genera. The sections of the group we adopt are the following:—

Sexes similar

Hylocichla. Smallest species. Bill short, broad at base; much depressed. Tarsi long and slender, longer than middle toe and claw, by the additional length of the claw; outstretched legs reaching nearly to tip of tail. Body slender. Color: above olivaceous or reddish, beneath whitish; breast spotted; throat without spots.

Turdus. Bill stouter and higher. Tarsi stout and short, scarcely longer than middle toe and claw. Body stout, generally whitish beneath and spotted. (Second quill longer than fifth?)

Planesticus. Similar to preceding. (Second quill shorter than fifth?) Beneath mostly uni-colored; unstreaked except the throat, which is whitish with dark streaks.

Sexes dissimilar

Merula. Similar to Turdus. Male usually more or less black, especially on the head; females brownish, often with streaked throats. Bill distinctly notched.

Hesperocichla. Similar to Turdus. Male reddish beneath, with a black collar. Bill without notch.

Subgenus HYLOCICHLA, Baird

Hylocichla, Baird (s. g.), Rev. Am. Birds, 1864, 12. (Type, Turdus mustelinus.)

Turdus mustelinus.

1570

The essential characters of Hylocichla have already been given. The subgenus includes the small North American species, with Turdus mustelinus, Gm., at the head as type, which are closely connected on the one side with Catharus, by their lengthened tarsi, and with Turdus by the shape of the wing. The bills are shorter, more depressed, and broader at base than in typical Turdus, so much so that the species have frequently been described under Muscicapa.

It is not at all improbable that naturalists may ultimately conclude to consider the group as of generic rank.

In this group there appears to be five well-marked forms or “species.” They are, mustelinus, Gm., pallasi, Caban., fuscescens, Steph., swainsoni, Caban., and aliciæ, Baird. The first-named is totally unlike the rest, which are more closely related in appearance.

In studying carefully a very large series of specimens of all the species, the following facts become evident:—

1. In autumn and winter the “olive” color of the plumage assumes a browner cast than at other seasons; this variation, however, is the same in all the species (and varieties), so that in autumn and winter the several species differ from each other as much as they do in spring and summer.

Turdus ustulatus.

Of these five species, two only (pallasi and swainsoni) inhabit the whole breadth of the continent; and they, in the three Faunal Provinces over which they extend, are modified into “races” or “varieties” characteristic of each region. The first of these species, as the pallasi var. pallasi, extends westward to the Rocky Mountains, and migrates in winter into the South; specimens are very much browner in the winter than in spring; but in the Rocky Mountain region is a larger, grayer race, the var. auduboni. This, in its migrations, extends along the central mountain region through Mexico to Guatemala; specimens from the northern and southern extremes of this range are identical in all the specific characters; but the southern specimens, being in the fall and winter dress, are browner in color than northern ones (spring birds); an autumnal example from Cantonment Burgwyn, N. M., is as brown as any Central American specimen. Along the Pacific Province, from Kodiak to Western Mexico, and occasionally straggling eastward toward the Rocky Mountain system, there is the var. nanus, a race smaller than the var. pallasi, and with much the same colors as var. auduboni, though the rufous of the tail is deeper than in either of the other forms. In this race, as in the others, there is no difference in size between specimens from north and south extremes of its distribution, because the breeding-place is in the North, all Southern specimens being winter sojourners from their Northern birthplace.