Natural History of the Mammalia of India and Ceylon

The stories of their attacking and killing tigers must be received with caution, though it is certain they will harass both tigers and leopards. I wrote some time back, in 'Seonee': "The natives in all parts of India declare that even tigers are attacked by them; and we once heard a very circumstantial account given of a fight, which took place near the station of Seonee, between a tiger and a pack of these dogs, in which the latter were victors. They followed him about cautiously, avoiding too close a contact, and worried him for three successive days—a statement which should be received with caution. We have, however, heard of them annoying a tiger to such an extent as to make him surrender to them the prey which he had killed for himself."

I agree with Jerdon in disbelieving the native superstition that the wild dog sheds a pungent secretion on his tail, and whisks it in the eyes of the animals it attacks, or covers the leaves of the bushes through which the victim graze, and then takes advantage of the temporary blindness thus caused; but it is a curious fact that the idea is prevalent in all parts of India, north and south, and has been accepted by many writers on Indian sports.

The wild dog dwells and breeds in holes and caves in rocks. The breeding season is from January to March, and about six whelps are born at a time. The mammæ are more numerous than in any other canine—from twelve to fourteen. Jerdon notices that Mr. Wilson at Simla discovered a breeding-place in holes under some rocks, where evidently several females were breeding together. At such times they endeavour to hunt their game towards their den, and kill it as near to it as possible.

GENUS VULPES

The foxes form a distinct group of the Canidæ; their bodies are long, with short legs, the muzzle more lengthened in comparison and much sharper, and the pupil of the eye contracts vertically instead of circularly; the tail is very bushy, with a gland at the base secreting a strong odorous substance. The female has six mammæ. There are two types in India—the desert fox or fox of the plains, Cynalopex of Hamilton Smith; and the hill fox, which approximates to the European species. The former has longer ears and longer and more slender limbs.

NO. 250. VULPES BENGALENSISThe Indian Fox (Jerdon's No. 138)

NATIVE NAMES.—Lomri, Lokri, Lokeria, Hindi; Kokri, Mahrathi; Khekar and Khikir in Behar; Khek-sial, Bengali; Konk, Kemp-nari, Chanaak-nari, Canarese; Konka-nakka or Gunta-nakka, Poti-nara, Telegu.—Jerdon.

HABITAT.—Throughout India; probably Ceylon, as Kellaart mentions having heard of a fox there, but I cannot trace it, or any other, in Burmah.

DESCRIPTION.—"Reddish-grey; rufous on the legs and muzzle; reddish white beneath; ears long dark brown externally; tail long bushy, with a broad black tip; muzzle very acute; chin and throat whitish."—Jerdon.

Here is Colonel Sykes's description of it in Southern India:—

"It is a very pretty animal, but smaller than the European fox; head short; muzzle very sharp; eyes oblique; irides nut-brown; legs very slender; tail trailing on the ground, very bushy; along the back and on the forehead fawn colour, with hair having a white ring to its tip; back, neck, between the eyes, along the sides, and half way down the tail reddish-grey; each hair banded black and reddish-white; all the legs reddish outside, reddish-white inside; chin and throat dirty white; along the belly reddish-white; ears externally dark brown, and with the fur so short as to be scarcely discoverable; edges of eyelids black; muzzle red brown."

The colour however varies a good deal, according to season and locality. It becomes more grey in the cold season. McMaster writes that he once killed one silvery grey, almost white.

SIZE.—Head and body, 20 to 21 inches; tail, 12 to 14 inches; weight, 5½ lbs.

This fox is common, not only in open country, but even in cantonments and suburbs of cities. Hardly a night passes without its familiar little chattering bark in the Dalhousie Square gardens, or on the Maidan, being heard; and few passengers running up and down our railway lines, who are on the look-out for birds and animals as the train whirls along, fail to see in the early morning our little grey friend sneaking home with his brush trailing behind him.

Jerdon says of the manner in which he carries this that he trails it when going slowly or hunting for food; holds it out horizontal when running; and raises it almost erect when making a sudden turn.

It also, like the jackal; will eat fruit, such as melons, ber, &c., and herbs. It breeds in the spring, from February to April, and has four cubs. Jerdon says the cubs are seldom to be seen outside their earth till nearly full grown. It is much coursed with greyhounds, and gives most amusing sport, doubling constantly till it gets near an earth; but it has little or no smell, so its scent does not lie.

Sir Walter Elliot wrote of it in the Madras Journal of Literature and Science (vol. x. p. 102): "Its principal food is rats, land-crabs, grasshoppers, beetles, &c. On one occasion a half-devoured mango was found in the stomach. It always burrows in open plains, runs with great speed, doubling like a hare; but instead of stretching out at first like that animal, and trusting to its turns as a last resource, the fox turns more at first; and, if it can fatigue the dogs, it then goes straight away."

It is easily tamed if taken young, and is very playful, but Jerdon, in repeating the assertion that tame foxes sooner or later go mad, says he has known one or two instances where they have done so; but McMaster throws doubt on this, and puts the supposed madness down to excitement at the amorous season. He gives an interesting account of a pair kept by a friend, which lived on amicable terms with his greyhounds. The owner writes: "I sometimes took them on to the parade ground, and slipped a couple of greyhounds after them. They never ran far, as when tired they lay down on their backs, and were at once recognised by the dogs. On one occasion one fox was tired before the other, and after he had made friends with the dogs he joined them in the chase after the other."

NO. 251. VULPES LEUCOPUSThe Desert Fox (Jerdon's No. 139)

HABITAT.—Northern India, and also on the Western Coast about Cutch.

DESCRIPTION.—"Light fulvous on the face, middle of back and upper part of tail; cheeks, sides of neck and body, inner side, and most of the fore parts of the limbs, white; shoulder and haunch, and outside of the limbs nearly to the middle joint, mixed black and white; tail darker at the base above, largely tipped with white; lower parts nigrescent; ears black posteriorly; fur soft and fine as in V. montanus, altogether dissimilar from that of V. Bengalensis. The skull with the muzzle distinctly narrower, and the lower jaw weaker. One I killed at Hissar had the upper parts fulvous, the hair black-tipped; sides paler; whole lower parts from the chin, including the inside of the arm and thigh, blackish; feet white on the inner side anteriorly, with a blackish border on the anterior limbs; legs fulvous externally; all feet white; tail always with a white tip."—Jerdon.

SIZE.—Head and body, 20 inches; tail, 14 inches; weight, 5½ lbs.

According to Mountstuart Elphinstone the backs of the foxes in Hurriana are of the same colour as the common fox, but in one part of the desert their legs and belly, up to a certain height, are black, and in another white—the one seems to have been wading up to the belly in ink, and the other in whitewash.

This fox lives chiefly on the jerboa-rat (Gerbillus Indicus) common on sandy plains. Jerdon thinks it more speedy than the common Indian fox.

NO. 252. VULPES FERRILATUSThe Thibetan Grey Fox

NATIVE NAME.—Iger, Thibetan.

HABITAT.—Thibet.

DESCRIPTION.—Pale fulvous, with grizzled white or iron-grey sides; shorter ears than in the Indian fox.

We now come to the true foxes, with shorter legs and moderate ears.

NO. 253. VULPES MONTANUSThe Hill Fox (Jerdon's No. 140)

NATIVE NAMES.—Loh, Kashmiri; Lomri, Hindi, at Simla; Wamu, Nepalese.

HABITAT.—Throughout the Himalayas.

DESCRIPTION.—Pale fulvous, with a dark brownish or deep chestnut streak down the back; sides deeper fulvous; the haunches a steely grey, mixed with yellowish hairs; tail grey and very bushy, largely tipped with white; ears deep black on outside; cheeks and jowl greyish-white; moustaches black; legs chestnut in front, paling off behind.

SIZE.—Head and body, 30 inches; tail, 19 inches; weight, 14 lbs.

Not at all unlike an English fox, only more variegated. The foregoing description is taken chiefly from a very fine specimen shot in the garden of the house in which I stayed at Simla; but it is subject to great variation, and is in its chief beauty in its winter dress. Several specimens which I have seen are all more or less different in colour. I have never seen a handsomer fox; the fur is extremely rich, the longer hairs exceeding two inches, and the inner fur is fine and dense. It is said to breed in April and May, the female usually having three to four cubs.

NO. 254. VULPES PUSILLUSThe Punjab Fox (Jerdon's No. 141)

HABITAT.—Punjab Salt Range.

DESCRIPTION.—Similar to the last, but much smaller, being about the size of the Indian fox. Jerdon suggests that it may be a variety of the last species, dwarfed by a warmer climate, but Blyth and others keep it apart.

NO. 255. VULPES FLAVESCENSThe Persian Fox

NATIVE NAMES.—Tulke, at Yarkand; Wamu, Nepalese.

HABITAT.—Eastern Turkestan, Ladakh, Persia, and, according to Gray, Indian Salt Range; Thibet.

DESCRIPTION.—Fulvous, darker on back, very similar to V. montanus, only more generally rufous and paler, with longer hair and larger teeth; face, outer side of fore-legs and base of tail pale fulvous; spot on side of face, chin, front of fore-legs, and a round spot on upper part of hind foot blackish; hairs of tail tipped black; ears externally black; tail tipped largely with white. The skull of one mentioned by Mr. Blanford had larger auditory bullæ than either the European fox or V. montanus.

NO. 256. VULPES GRIFFITHIIThe Afghanistan Fox

This was at first reckoned by Blyth as synonymous with the last, but was afterwards separated and renamed. It is stated by Hutton to be common about Candahar, where the skins are made into reemchas and poshteens, the price in 1845 being about six annas a skin.

MARINE CARNIVORA

We disposed of the land Carnivora in the last article, and now, before proceeding to the Cetacea, I will give a slight sketch of the marine Carnivora, of which, however, no examples are to be found on the Indian coasts. The Pinnipedia or Pinnigrada are amphibious in their habits, living chiefly in the water, but resorting occasionally to the land. There are some examples of the land Carnivora which do the same—the polar bear and otter, and more especially the sea-otter, Enhydra lutris, which is almost exclusively aquatic, but these are all decidedly of the quadrupedal type, whereas in the amphibia we see the approach to the fish form necessary for their mode of life. The skeleton reveals the ordinary characteristics of the quadruped with somewhat distorted limbs. The bones of the forelimbs are very powerful and short, a broad scapula, short humerus and the ulna and radius are stout, parallel to each other, and the latter much broader at the base; often in old animals the two are ankylosed at the joint, which is also the case with the tibia and fibula. The hip-bones are narrow and much compressed, the femur remarkably short, the shank-bones and the bones of the feet very long. In walking on land the feet are, in the case of the Otaria or eared seals placed flat on the full sole; the common seals never use their hind limbs on the shore. The dentition is essentially carnivorous, but varies considerably in the different families, and even in the Phocidæ themselves. The stomach is simple, but the intestines are considerably longer than in the Felidæ, averaging about fifteen times the length of the body; the digestion is rapid. The bones are light and spongy, and the spine particularly flexible, from the amount of cartilage between the bones. They have a large venous cavity in the liver, and the lungs are capacious, the two combining to assist them in keeping under water; the blood is dark and abundant. The brain is large, and in quantity and amount of convolution exceeds that of the land Carnivores. Their hearing is acute, but their sight out of water is defective.

Their external features are an elongated pisciform body, the toes joined by a membrane converting the feet into broad flippers or fins, the two hind ones being so close as to act like the caudal fin of a fish. The head is flattish and elongated, or more or less rounded, but in comparison with the body it is small. Except in the Otaridæ there are no perceptible ears, and in them the ear is very small. The fur is of two kinds, one long and coarse, but the other, or under fur, is beautifully soft and close, and is the ordinary sealskin of commerce. The roots of the coarse hair go deeper into the skin than those of the under fur, so the furrier takes advantage of this by thinning the skin down to the coarse roots, cutting them free, and then the hairs are easily removed, leaving the soft fur attached to the skin.

The Pinnigrada are divided into three families—the Trichechidæ, or walruses; the Otaridæ, or sea-lions or eared seals; and the Phocidæ, or ordinary seals.

As none of these animals have been as yet observed in the Indian seas, being chiefly denizens of cold zones, I will not attempt any further description of species, having merely alluded to them en passant as forming an important link in the chain of animal creation.

We must now pass on to the next order, a still more aquatic one.

ORDER CETACEA—THE WHALES

These curious creatures have nothing of the fish about them, save the form, and frequently the name. In other respects they are warm-blooded, viviparous mammals, destitute of hinder limbs, and with very short fore-limbs completely enclosed in skin, but having the usual number of bones, though very much shortened, forming a kind of fin. The fin on the back is horizontal, and not rayed and upright like that of a fish; the tail resembles that of a fish in form, the caudal vertebræ running through the middle of it. The immense muscular power of this tail, with its broad flanges, arises from the flesh of the body, terminating in long cords of tendon, running to the tip. The vertebral column is often ankylosed in the fore-part, but is extremely elastic, owing to the cartilaginous cushion between each bone in the latter half. Thus, whilst the fore-part is rigid, the hinder is flexible in the extreme. The brain is large and much convoluted; the heart is very large, and the blood-vessels extremely full and numerous, with extensive ramifications, which, being filled with oxygenated blood, assist in supporting life whilst submerged. The lungs are also very large. The laryngeal and nasal passages are peculiar. The following description is by Dr. Murie: "In front of the larynx of man we all know that there is an elastic lid, the epiglottis, which folds over and protects the air passage as food is swallowed. The side cartilages constitute the walls of the organ of voice and protect the vocal chords. Now, in the comparatively voiceless whale, the cartilages, including the epiglottis, form a long rigid cylindrical tube, which is thrust up the passage at the back of the palate in continuity with the blow-hole. It is there held in place by a muscular ring. With the larynx thus retained bolt upright, and the blow-hole being meanwhile compressed or closed, the cetacean is enabled to swallow food under water without the latter entering the lungs." The stomach is peculiar, being composed of several sacs or chambers with narrow passages between; the intestines are long, glandular and, according to Dr. Murie, full of little pouches. There is no gall bladder; the gullet is very narrow in some and wider in others. Some have teeth, others are without. The eyes are small; the ears deficient externally, though the interior small ear-bones of ordinary mammals are in these massive and exceedingly dense, so much so, as Murie observes, as to be frequently preserved fossil when other osseous structures are destroyed.

The cetacea have been divided into the Denticete, or Toothed Whales, and the Mysticete, or Whalebone Whales. The former contains the river dolphins, the ziphoid whales, the gigantic sperm whale, the sea dolphins, and the narwhal or sea unicorn. The latter contains the baleen whales.

DENTICETE—THE TOOTHED WHALES

None of the larger species are found on these coasts, or in the Indian Ocean, the two most interesting of which are the gigantic sperm whales (Physeter macrocephalus), and the curious narwhal or sea unicorn (Monodon monoceros). The latter is an inhabitant of the northern seas only, but the sperm abounds in warmer waters, being frequently found in the sub-tropical oceans. I have occasionally seen them in the South Atlantic, though they are said to have diminished there of late years. It is a wonder that the species does not get scarce in many localities, so great is the chase after them. During the last forty years the Americans alone have taken at the rate of 10,000 barrels of sperm oil per annum, or upwards of four million barrels since 1835. The sperm whale, though of such enormous bulk and courage, yet has enemies besides man. The thrasher and the killer whale both attack it, and sailors assert that the sword-fish and thrasher combine against it, the latter stabbing from below, whilst the former leaps on it with stunning blows. I think by sword-fish (Xiphias), which is also a large but not so very sanguinary a fish, they mean the saw-fish (Pristis), which is allied to the sharks, and which attacks the largest whales. The sword-fish has however the character of being pugnacious. The old sperms, especially males, will show fight at times, but the younger ones are easily alarmed, and on being molested rush off in various directions, each looking out for himself. The sperm whale is known from the others by the way in which it spouts, the jet being thrown up obliquely forwards, and it blows at regular intervals. Although the old "bulls" show a certain amount of ferocity at times, their savageness is considerably exaggerated by the whalers, who love to spin yarns about them. Having watched the habits of these and the baleen whales with curiosity, I tried to get as much information about them as I could, from the whalers, but, with the exception of the officers of whaling ships, there was much that was unreliable in Jack's notions about the sperm. On one occasion I was just too late to see one killed. The boats, under full sail, were towing the carcase towards the ship. I would have given a good deal to have seen the encounter. The food of the sperm consists greatly of the huge rock squid or cuttle-fish, which they swallow in large lumps. I have heard whalers assert that a wounded sperm in the death agony will vomit immense pieces of squid. In this respect it differs much from the baleen whales, which have a narrow gullet. According to Professor Flower there is no sufficient evidence of the existence of more than one species of sperm whales, but an allied species, Physeter (Euphysetes) simus, is found on the Madras coast, and to this I will allude further on.

FAMILY DELPHINIDÆ—THE DOLPHINS OR PORPOISESGENUS PLATANISTA—THE RIVER DOLPHINS

A globular head with a long, compressed and, towards the end, spoon-shaped rostrum or snout; flippers short, broad and triangular; a long body of moderate girth; no back fin, but a slight elevation which takes its place. There is a decided depression between the head and body on the region of the neck; the eye is remarkably small, so much so as to be hardly perceptible; in an adult of eight feet long the whole eye-ball is no bigger than a pea, and the orifice of the ear is like a pin-hole.

The skull has peculiar features. "The apparently rounded skull behind the snout has broad, thick zygomatic arches, and above and in front of these the cheek-bones (maxillæ) each send forwards and inwards a great roughened sheet of bone or crest, which forms a kind of open helmet. In the large hollow between these bony plates, and somewhat behind, are situated the nasal orifices, which are slightly awry" (Murie).19 Professor Flower's notice of the skull ('Osteology of the Mammalia') is thus worded: "The orbit is extremely small, the temporal fossa large, and the zygomatic processes of the squamosal are greatly developed. From the outer edge of the ascending plates of the maxillæ, which lie over the frontals, great crests of bone, smooth externally, but reticulated and laminated on their inner surface, rise upwards, and, curving inwards, nearly meet in the middle line above the upper part of the face."

The dentition is also curious, the upper and lower jaws being provided with a number of teeth, pointed and conical in front, and smaller and more flattened behind. They vary in number. In an example quoted by Dr. Murie the total was 117, viz., 27—28/30—32, but in a specimen examined by Dr. Anderson, who has most exhaustively described these animals, the total number of teeth amounted to 128, i.e. 33—32/32—31. (See Appendix B.)

The cervical vertebræ are movable, and not ankylosed, as in many of the cetacea; the cæcum is small; the blow-hole is a narrow slit, not transverse as in other whales, but longitudinal. I have somewhat gone out of order in Jerdon's numbering in bringing in this genus here instead of letting it follow Delphinus, as he has done. These river Dolphins naturally come after the extinct Phocodontia or seal-toothed whales, and bear considerable resemblance in the dentition to the extinct genus Squalodon.

NO. 257. PLATANISTA GANGETICAThe Gangetic Porpoise (Jerdon's Nos. 144 and 145)

NATIVE NAMES.—Soonse, Soosoo, Soosa, Hindi; Susak, Shishuk, Bengali; Sisumar, Sanscrit; Bulhan or Sunsar, on the Indus; Hihoo, Siho; Assamese; Huhh in Cachar and Sylhet.

HABITAT.—In the larger rivers connected with the Ganges nearly up to the hills; also in the Brahmaputra and in the Indus, but in fresh water; only it does not go out to sea.

DESCRIPTION.—"A long compressed snout with a formidable array of teeth; a vaulted compressed forehead; longitudinal blow-hole; scarcely perceptible eye; distinct neck; broad and abruptly truncated pectoral fins, and small dorsal fin; and the male, a smaller but heavier-built animal than the female, with a shorter snout" (Anderson). The colour is from a dark lead to a sooty black; according to Jerdon "when old with some lighter spots here and there; shining pearl-grey when dry."

SIZE.—From six to eight feet.

This animal, though not often captured, at all events in the vicinity of Calcutta, is familiar to most people who have travelled on the larger Indian rivers. It is common enough in the Hooghly. I have frequently observed it in the river abreast of the Fort whilst we were slowly driving down the Course.

I am largely indebted to Dr. Anderson for information concerning it, for he has not only most carefully watched the habits of this curious animal, but has most exhaustively described its anatomy in his 'Anatomical and Zoological Researches.' It is found in the Hooghly, chiefly in the cold weather, migrating during the hot and rainy season; at least so it was supposed, and Dr. Cantor conjectured that at such times it visited the sea, but this has been proved to be not the case. The soosoo never leaves fresh water; and it is in the river during the rains, for fishermen catch it in their nets, but it is hardly ever seen at that time. It rises so as to expose the blow-hole only, and the rush of the swollen waters prevents the peculiar sound of respiration being heard. But in the cold weather, when the river is calm, the ear is attracted at once by the hissing puff of expiration, and the animal may be seen to bound almost out of the water. Dr. Anderson had one alive in captivity for ten days, and carefully watched its respirations. "The blow-hole opened whenever it reached the surface of the water. The characteristic expiratory sound was produced, and so rapid was the inspiration that the blow-hole seemed to close immediately after the expiratory act." He states that "the respirations were tolerably frequent, occurring at intervals of about one-half or three-quarters of a minute, and the whole act did not take more than a few seconds for its fulfilment." But it is probable that in a free state and in perfect health the animal remains longer under water. It has certainly been longer on several occasions when I have watched for the reappearance of one in the river. The food of the Gangetic dolphin consists chiefly of fish and crustacea; occasionally grains of rice and remains of insects are found in the stomach, but these are doubtless, as Dr. Anderson conjectures, in the fish swallowed by the dolphin. The period of gestation is said to be eight to nine months, and usually only one at a time is born, between April and July. The young are sometimes caught with their mothers, and are said to cling by holding on by the mouth to the base of the parent's pectoral fins. "The flesh and blubber are occasionally eaten by many of the low caste Hindus of India, such as the Gurhwals, the Domes of Jessore and Dacca districts, the Harrees, Bourees, Bunos, Bunpurs, Tekas, Tollahas, the Domes of Burdwan and Bhagulpore, who compare it to venison; also by the Teewars and Machooas of Patna, the Mussahars of Shahabad, the Gourhs and Teers of Tirhoot, and the Mullahs of Sarun. In the North-west Provinces about Allahabad, the Chumars, Passees, Kooras, Khewuts or Mullahs, have rather a high estimate of the flesh, which they assert resembles turtle. The Koonths of Benares, Phunkeahs, Natehmurrahs, and Buahoas of Moradabad, and also such gipsy tribes as the Sainsees, Kunjars and Hubbossahs, in the neighbourhood of Meerut, do not despise it. In the Punjab we find the Choorahs, Dhapels, Sainsees, Budous, and Burars eating the flesh; and in Sind the Kehuls. The Moras, a tribe of Mahomedan boatmen who lead a wandering life on the streams in the Punjab and in Sind, subsist on the dolphin when by good chance they catch one; this is also the case with the Cacharies and the Nagas of Assam. The Sansee women on the Indus eat the flesh under the idea that it makes them prolific. All along the Ganges, Brahmahputra, and Indus, the oil is universally considered as of great value as an embrocation in rheumatism and for giving much strength when rubbed on the back and loins. But many other animal oils, such as those of various species of turtle, the crocodile, and the pelican, have a similar reputation. It is said to be of a very penetrating nature, and, owing to this property, it is highly prized for preserving leather, such as harness, &c. The illuminating powers of this oil are said to be very high." (Anderson's 'Anatomical and Zoological Researches.')