Natural History of the Mammalia of India and Ceylon

SIZE.—About same as H. hooluck.

It is, however, found in every variety of colour, from black to brownish, and variegated with light-coloured patches, and occasionally of a fulvous white. For a long time I supposed it to be synonymous with H. agilis of Cuvier, or H. variegatus of Temminck, but both Mr. Blyth and Dr. Anderson separate it. Blyth mentions a significant fact in distinguishing the two Indian Gibbons, whatever be their variations of colour, viz.: "H. hooluck has constantly a broad white frontal band either continuous or divided in the middle, while H. lar has invariably white hands and feet, less brightly so in some, and a white ring encircling the visage, which is seldom incomplete."3

H. lar has sometimes the index and middle fingers connected by a web, as in the case of H. syndactylus (a Sumatran species very distinct in other respects). The very closely allied H. agilis has also this peculiarity in occasional specimens. This Gibbon was called "agilis" by Cuvier from its extreme rapidity in springing from branch to branch. Duvaucel says: "The velocity of its movements is wonderful; it escapes like a bird on the wing. Ascending rapidly to the top of a tree, it then seizes a flexible branch, swings itself two or three times to gain the necessary impetus, and then launches itself forward, repeatedly clearing in succession, without effort and without fatigue, spaces of forty feet."

Sir Stamford Raffles writes that it is believed in Sumatra that it is so jealous that if in captivity preference be given to one over another, the neglected one will die of grief; and he found that one he had sickened under similar circumstances and did not recover till his rival (a Siamang, H. syndactylus) was removed.

NO. 3. HYLOBATES SYNDACTYLUSThe Siamang

HABITAT.—Tenasserim Province, Sumatra, Malayan Peninsula.

DESCRIPTION.—A more robust and thick-set animal than the two last; deep, woolly, black fur; no white supercilium nor white round the face. The skull is distinguished from the skull of the other Gibbons, according to Dr. Anderson, by the greater forward projection of the supraorbital ridges, and by its much deeper face, and the occipital region more abruptly truncated than in the other species. The index and middle toes of the foot are united to the last phalange.

SIZE.—About three feet.

This Gibbon is included in the Indian group on the authority of Helfer, who stated it to be found in the southern parts of the Tenasserim province. Blyth mentions another distinguishing characteristic—it is not only larger than the other Gibbons, but it possesses an inflatable laryngeal sac. Its arms are immense—five feet across in an adult of three feet high.

The other species of this genus inhabiting adjacent and other countries are H. Pileatus and H. leucogenys in Siam; H. leuciscus, Java; H. Mulleri and H. concolor, Borneo.

GENUS PRESBYTES—CUVIER'S GENUS SEMNOPITHECUS

These monkeys are characterised by their slender bodies and long limbs and tails. Jerdon says the Germans call them Slim-apes. Other striking peculiarities are the absence of cheek pouches, which, if present, are but rudimentary. Then they differ from the true monkeys (Cercopithecus) by the form of the last molar tooth in the lower jaw, which has five tubercles instead of four; and, finally, they are to be distinguished by the peculiar structure of the stomach, which is singularly complicated, almost as much so as in the case of Ruminants, which have four divisions. The stomach of this genus of monkey consists of three divisions: 1st, a simple cardiac pouch with smooth parietes; 2nd, a wide sacculated middle portion; 3rd, a narrow elongated canal, sacculated at first, and of simple structure towards the termination. Cuvier from this supposes it to be more herbivorous than other genera, and considers this conclusion justified by the blunter tubercles of the molars and greater length of intestines and cæcum, all of which point to a vegetable diet. "The head is round, the face but little produced, having a high facial angle."—Jerdon.

But the tout ensemble of the Langur is so peculiar that no one who has once been told of a long, loosed-limbed, slender monkey with a prodigious tail, black face, with overhanging brows of long stiff black hair, projecting like a pent-house, would fail to recognise the animal.

The Hanuman monkey is reverenced by the Hindus. Hanuman was the son of Pavana, god of the winds; his strength was enormous, but in attempting to seize the sun he was struck by Indra with a thunderbolt which broke his jaw (hanu), whereupon his father shut himself up in a cave, and would not let a breeze cool the earth till the gods had promised his son immortality. Hanuman aided Rama in his attack upon Ceylon, and by his superhuman strength mountains were torn up and cast into the sea, so as to form a bridge of rocks across the Straits of Manar.4

The species of this genus of monkey abound throughout the Peninsula. All Indian sportsmen are familiar with their habits, and have often been assisted by them in tracking a tiger. Their loud whoops and immense bounds from tree to tree when excited, or the flashing of their white teeth as they gibber at their lurking foe, have often told the shikari of the whereabouts of the object of his search. The Langurs take enormous leaps, twenty-five feet in width, with thirty to forty in a drop, and never miss a branch. I have watched them often in the Central Indian jungles. Emerson Tennent graphically describes this: "When disturbed their leaps are prodigious, but generally speaking their progress is not made so much by leaping as by swinging from branch to branch, using their powerful arms alternately, and, when baffled by distance, flinging themselves obliquely so as to catch the lower boughs of an opposite tree, the momentum acquired by their descent being sufficient to cause a rebound of the branch that carries them upwards again till they can grasp a higher and more distant one, and thus continue their headlong flight."

Jerdon's statement that they can run with great rapidity on all-fours is qualified by McMaster, who easily ran down a large male on horseback on getting him out on a plain.

A correspondent of the Asian, quoting from the Indian Medical Gazette for 1870, states that experiments with one of this genus (Presbytes entellus) showed that strychnine has no effect on Langurs—as much as five grains were given within an hour without effect. "From a quarter to half of a grain will kill a dog in from five to ten minutes, and even one twenty-fourth of a grain will have a decided tetanic effect in human beings of delicate temperament."—Cooley's Cycl. Two days after ten grains of strychnine were dissolved in spirits of wine, and mixed with rum and water, cold but sweet, which the animal drank with relish, and remained unhurt.

The same experiment was tried with one of another genus (Inuus rhesus), who rejected the poisoned fruit at once, and on having strychnine in solution poured down his throat, died.

The Langur was then tried with cyanide of potassium, which he rejected at once, but on being forced to take a few grains, was dead in a few seconds.

Although we may not sympathize with those who practise such cruel experiments as these above alluded to, the facts elucidated are worth recording, and tend to prove the peculiar herbivorous nature of this genus, which, in common with other strictly herbivorous animals, instinctively knows what to choose and what to avoid, and can partake, without danger, of some of the most virulent vegetable poisons. It is possible that in the forests they eat the fruit of the Strychnos nux-vomica, which is also the favourite food of the pied hornbill (Hydrocissa coronata).

NO. 4. SEMNOPITHECUS vel PRESBYTES ENTELLUSThe Bengal Langur (Jerdon's No. 1)

NATIVE NAMES.—Langur, Hanuman, Hindi; Wanur and Makur, Mahratti; Musya, Canarese.

HABITAT.—Bengal and Central India.

DESCRIPTION.—Pale dirty or ashy grey; darker on the shoulders and rump; greyish-brown on the tail; paler on the head and lower parts; hands and feet black.

SIZE.—Length of male thirty inches to root of tail; tail forty-three inches.

The Entellus monkey is in some parts of India deemed sacred, and is permitted by the Hindus to plunder their grain-shops with impunity; but I think that with increasing hard times the Hanumans are not allowed such freedom as they used to have, and in most parts of India I have been in they are considered an unmitigated nuisance, and the people have implored the aid of Europeans to get rid of their tormentors. In the forest the Langur lives on grain, fruit, the pods of leguminous trees, and young buds and leaves. Sir Emerson Tennent notices the fondness of an allied species for the flowers of the red hibiscus (H. rosa sinensis). The female has usually only one young one, though sometimes twins. The very young babies have not black but light-coloured faces, which darken afterwards. I have always found them most difficult to rear, requiring almost as much attention as a human baby. Their diet and hours of feeding must be as systematically arranged; and if cow's milk be given it must be freely diluted with water—two-thirds to one-third milk when very young, and afterwards decreased to one-half. They are extremely susceptible to cold. In confinement they are quiet and gentle whilst young, but the old males are generally sullen and treacherous. Jerdon says, on the authority of the Bengal Sporting Magazine (August 1836), that the males live apart from the females, who have only one or two old males with each colony, and that they have fights at certain seasons, when the vanquished males receive charge of all the young ones of their own sex, with whom they retire to some neighbouring jungle. Blyth notices that in one locality he found only males of all ages, and in another chiefly females. I have found these monkeys mostly on the banks of streams in the forests of the Central Provinces; in fact, the presence of them anywhere in arid jungles is a sign that water is somewhere in the vicinity. They are timid creatures, and I have never seen the slightest disposition about them to show fight, whereas I was once most deliberately charged by the old males of a party of Rhesus monkeys. I was at the time on field service during the Mutiny, and, seeing several nursing mothers in the party, tried to run them down in the open and secure a baby; but they were too quick for me, and, on being attacked by the old males, I had to pistol the leader.

NO. 5. SEMNOPITHECUS vel PRESBYTES SCHISTACEUS.5The Himalayan Langur (Jerdon's No. 2)

NATIVE NAMES.—Langur, Hindi; Kamba Suhú, Lepcha; Kubup, Bhotia.

HABITAT.—The whole range of the Himalayas from Nepal to beyond Simla.

DESCRIPTION (after Hodgson).—Dark slaty above; head and lower parts pale yellowish; hands concolorous with body, or only a little darker; tail slightly tufted; hair on the crown of the head short and radiated; on the cheeks long, directed backwards, and covering the ears. Hutton's description is, dark greyish, with pale hands and feet, white head, dark face, white throat and breast, and white tip to the tail.

SIZE.—About thirty inches; tail, thirty-six inches.

Captain Hutton, writing from Mussoorie, says: "On the Simla side I observed them also, leaping and playing about, while the fir-trees, among which they sported, were loaded with snow-wreaths, at an elevation of 11,000 feet."—'Jour. As. Soc. Beng.' xiii. p. 471.

Dr. Anderson remarks on the skull of this species, that it can be easily distinguished from entellus by its larger size, the supraorbital ridge being less forwardly projected, and not forming so thick and wide a pent roof, but the most marked difference lies in the much longer facial portion of schistaceus; the teeth are also larger; the symphysis or junction of the lower jaw is considerably longer and broader, and the lower jaw itself is generally more massive and deep.

NO. 6. SEMNOPITHECUS vel PRESBYTES PRIAMUSThe Madras Langur

NATIVE NAME.—Gandangi, Telugu.

HABITAT.—The Coromandel Coast and Ceylon.

DESCRIPTION.—Ashy grey, with a pale reddish or chocolat-au-lait tint overlying the whole back and head; sides of the head, chin, throat, and beneath pale yellowish; hands and feet whitish; face, palms and fingers, and soles of feet and toes black; hair long and straight, not wavy; tail of the colour of the darker portion of the back, ending in a whitish tuft.—Jerdon.

SIZE.—About the same as P. entellus.

Blyth, who is followed by Jerdon, describes this monkey as having a compressed high vertical crest, but Dr. Anderson found that the specimens in the Indian Museum owed these crests to bad stuffing. Kellaart, however, mentions it, and calls the animal "the Crested Monkey." In Sir Emerson Tennent's figure of P. priamus a slight crest is noticeable; but Kellaart is very positive on this point, saying: "P. priamus is easily distinguished from all other known species of monkeys in Ceylon by its high compressed vertical crest."

Jerdon says this species is not found on the Malabar Coast, but neither he nor McMaster give much information regarding it. Emerson Tennent writes: "At Jaffna, and in other parts of the island where the population is comparatively numerous, these monkeys become so familiarised with the presence of man as to exhibit the utmost daring and indifference. A flock of them will take possession of a palmyra palm, and so effectually can they crouch and conceal themselves among the leaves that, on the slightest alarm, the whole party becomes invisible in an instant. The presence of a dog, however, excites such irrepressible curiosity that, in order to watch his movements, they never fail to betray themselves. They may be frequently seen congregated on the roof of a native hut; and, some years ago, the child of a European clergyman, stationed near Jaffna, having been left on the ground by the nurse, was so teased and bitten by them as to cause its death."

In these particulars this species resembles P. entellus.

NO. 7. SEMNOPITHECUS vel PRESBYTES JOHNIIThe Malabar Langur (Jerdon's No. 4)

HABITAT.—The Malabar Coast, from N. Lat. 14° or 15° to Cape Comorin.

DESCRIPTION.—Above dusky brown, slightly paling on the sides; crown, occiput, sides of head and beard fulvous, darkest on the crown; limbs and tail dark brown, almost black; beneath yellowish white.—Jerdon.

SIZE.—Not quite so large as P. entellus.

This monkey was named after a member of the Danish factory at Tranquebar, M. John, who first described it. It abounds in forests, and does not frequent villages, though it will visit gardens and fields, where, however, it shuns observation.

The young are of a sooty brown, or nearly black, without any indication of the light-coloured hood of the adult.

NO. 8. SEMNOPITHECUS vel PRESBYTES JUBATUSThe Nilgheri Langur (Jerdon's No. 5)

HABITAT.—The Nilgheri Hills, the Animallies, Pulneys, the Wynaad, and all the higher parts of the range of the Ghâts as low as Travancore.

DESCRIPTION.—Dark glossy black throughout, except head and nape, which are reddish brown; hair very long; in old individuals a greyish patch on the rump.—Jerdon.

SIZE.—Length of head and body, 26 inches; tail, 30.

This monkey does not, as a rule, descend lower than 2,500 to 3,000 feet; it is shy and wary. The fur is fine and glossy, and is much prized (Jerdon). Its flesh is excellent food for dogs (McMaster).

Dr. Anderson makes this synonymous with the last.

NO. 9. SEMNOPITHECUS vel PRESBYTES PILEATUSThe Capped Langur

HABITAT.—Assam, Chittagong, Tipperah.

DESCRIPTION.—General colour dark ashy grey, with a slight ferruginous tint; darker near head and on shoulders; underneath and on the inside of the limbs pale yellowish, with a darker shade of orange or golden yellow on the breast and belly. The crown of the head is densely covered with bristly hairs, regularly disposed and somewhat elongated on the vertex so as to resemble a cap, whence the name. Along the forehead is a superciliary crest of long black bristles, directed outwardly; whiskers full and down to the chin: behind the ears is a small tuft of white hairs; the tail is long, one third longer than the body, darker near the end, and tufted; fingers and toes black.

SIZE.—A little smaller than P. entellus.

This monkey is found in Northern Assam, Tipperah and southwards to Tenasserim; in Blyth's 'Catalogue of the Mammals of Burmah' it is mentioned as P. chrysogaster (Semnopithecus potenziani of Bonaparte and Peters). He writes of it: "Females and young have the lower parts white, or but faintly tinted with ferruginous, and the rest of the coat is of a pure grey; the face black, and there is no crest, but the hairs of the crown are so disposed as to appear like a small flat cap laid upon the top of the head. The old males seem always to be of a deep rust-colour on the cheeks, lower parts, and more or less on the outer side of the limbs; while in old females this rust colour is diluted or little more than indicated."

Dr. Anderson says that a young one he had was of a mild disposition, which however is not the character of the adult animal, which is uncertain, and the males when irritated are fierce, and determined in attack. No rule, however, is without its exception, for one adult male, possessed by Blyth, is reported as having been an exceeding gentle animal.

NO. 10. SEMNOPITHECUS vel PRESBYTES BARBEIThe Tipperah Langur

HABITAT.—Tipperah, Tenasserim.

DESCRIPTION.—No vertical crest of hair on the head, nor is the occipital hair directed downwards, as in the next species. Shoulders and outside of arm silvered; tail slightly paler than body, "which is of a blackish fuliginous hue."

More information is required about this monkey, which was named by Blyth after its donor to the Asiatic Society, the Rev. J. Barbe. Blyth considered it as distinct from P. Phayrei and P. obscurus, which last is from Malacca.

Dr. Anderson noticed it in the valley of the Tapeng in the centre of the Kakhyen Hills, in troops of thirty to fifty, in high forest trees overhanging the mountain streams. Being seldom disturbed, they permitted a near approach.

NO. 11. SEMNOPITHECUS vel PRESBYTES PHAYREI Syn.—SEMNOPITHECUS CRISTATUSThe Silvery-Leaf Monkey (Blyth)

HABITAT.—Arracan, Malayan Peninsula, Sumatra, Borneo.

DESCRIPTION.-Colour dusky grey-brown above, more or less dark, with black hands and feet; a conspicuous crest on the vertex; under parts white, scarcely extending to the inside of the limbs; sides grey like the back; whiskers dark, very long, concealing the ears in front; lips and eyelids conspicuously white, with white moustachial hairs above and similar hairs below.

SIZE.—Two feet; tail, 2 feet 6 inches.

This monkey was named by Blyth after Captain (now Sir Arthur) Phayre, who first brought it to his notice; but he afterwards reconciled it as being synonymous with Semnopithecus cristatus. The colouring, according to different authors, seems to vary considerably, which causes some confusion in description. It differs from an allied species, S. maurus, in selecting low marshy situations near the banks of streams. Its favourite food is the fruit of the Nibong palm (Oncosperma filamentosa).

NO. 12. SEMNOPITHECUS vel PRESBYTES OBSCURUSThe Dusky-Leaf Monkey

HABITAT.—Mergui and the Malayan Peninsula.

DESCRIPTION.—Adults ashy or brownish black, darker on forehead, sides of face, shoulder, and sides of body; the hair on the nape is lengthened and whitish. The newly-born young are of a golden ferruginous colour, which afterward changes to dusky-ash colour, the terminal half of the tail being last to change; the mouth and eyelids are whitish, but the rest of the face black.

SIZE.—Body, 1 foot 9 inches; tail, 2 feet 8 inches.

This monkey is most common in the Malayan Peninsula, but has been found to extend to Mergui, where Blyth states it was procured by the late Major Berdmore. Dr. Anderson says it is not unfrequently offered for sale in the Singapore market.

NO. 13. SEMNOPITHECUS vel PRESBYTES CEPHALOPTERUSThe Ceylon Langur

NATIVE NAME.—Kallu Wanderu.

HABITAT.—The low lands of Ceylon.

DESCRIPTION.—General colour cinereous black; croup and inside of thighs whitish; head rufescent brown; hair on crown short, semi-erect; occipital hairs long, albescent; whiskers white, thick and long, terminating at the chin in a short beard, and laterally angularly pointed; upper lip thinly fringed with white hairs; superciliary hairs black, long, stiff and standing erect; tail albescent and terminating in a beard tuft; face, palms, soles, fingers, toes and callosities black; irides brown.—Kellaart.

SIZE.—Length, 20 inches; tail 24 inches.

Sir E. Tennent says of this monkey that it is never found at a higher elevation than 1,300 feet (when it is replaced by the next species).

"It is an active and intelligent creature, little larger than the common bonneted macaque, and far from being so mischievous as others of the monkeys in the island. In captivity it is remarkable for the gravity of its demeanour and for an air of melancholy in its expression and movements, which are completely in character with its snowy beard and venerable aspect. In disposition it is gentle and confiding, sensible in the highest degree of kindness, and eager for endearing attention, uttering a low plaintive cry when its sympathies are excited. It is particularly cleanly in its habits when domesticated, and spends much of its time in trimming its fur and carefully divesting its hair of particles of dust. Those which I kept at my house near Colombo were chiefly fed upon plantains and bananas, but for nothing did they evince a greater partiality than the rose-coloured flowers of the red hibiscus (H. rosa sinensis). These they devoured with unequivocal gusto; they likewise relished the leaves of many other trees, and even the bark of a few of the more succulent ones."

NO. 14. SEMNOPITHECUS vel PRESBYTES URSINUSThe Great Wanderu

NATIVE NAME.—Maha Wanderu.

HABITAT.—The mountainous district of Ceylon.

DESCRIPTION.—Fur long, almost uniformly greyish black; whiskers full and white; occiput and croup in old specimens paler coloured; hands and feet blackish; tail long, getting lighter towards the lower half. The young and adults under middle age have a rufous tint, corresponding with that of the head of all ages.

SIZE.—Body about 22 inches; tail, 26 inches.

The name Wanderu is a corruption of the Singhalese generic word for monkey, Ouandura, or Wandura, which bears a striking resemblance to the Hindi Bandra, commonly called Bandar—b and v being interchangeable—and is evidently derived from the Sanscrit Banur, which in the south again becomes Wanur, and further south, in Ceylon, Wandura. There has been a certain amount of confusion between this animal and Inuus silenus, the lion monkey, which had the name Wanderu applied to it by Buffon, and it is so figured in Cuvier. They are both large monkeys, with great beards of light coloured hair, but in no other respect do they resemble. Sir Emerson Tennent says: "It is rarely seen by Europeans, this portion of the country having till very recently been but partially opened; and even now it is difficult to observe its habits, as it seldom approaches the few roads which wind through these deep solitudes. At early morning, ere the day begins to dawn, its loud and peculiar howl, which consists of quick repetition of the sound how-how! may be frequently heard in the mountain jungles, and forms one of the characteristic noises of these lofty situations." This was written in 1861; since then much of the mountainous forest land has been cleared for coffee-planting, and the Wanderu either driven into corners or become more familiarised with man. More therefore must be known of its habits by this time, and information regarding it is desirable.