Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras

"It must not, however, be supposed that the southern animals migrated from the Mediterranean area as far north as Yorkshire in the same year, or the northern as far south as the Mediterranean. There were, as we shall see presently, secular changes of climate in Pleistocene Europe, and while the cold was at its maximum the arctic animals arrived at the southern limit, and while it was at its minimum the spotted hyæna and hippopotamus and other southern animals roamed to their northern limit. Thus every part of the middle zone has been successively the frontier between the northern and southern groups, and consequently their remains are mingled together in the caverns and river-deposits, under conditions which prove them to have been contemporaries in the same region. In some of the caverns, such as that of Kirkdale, the hyæna preyed upon the reindeer at one time of the year and the hippopotamus at another. In this manner the association of northern and southern animals may be explained by their migration according to the seasons; and their association over so wide an area as the middle zone, by the secular changes of climate by which each part of the zone in turn was traversed by the advancing and retreating animals."41

When we consider that remains of the hippopotamus have been found in the caves of North Wales and Bristol as well as in those of Yorkshire, associated in all with the reindeer and in some with the woolly rhinoceros or the mammoth, and that the animal must have reached these localities by means of slow-flowing rivers or flooded marshes by very circuitous routes, we shall be convinced that these long journeys from the warmer regions of South Europe could not have been made during the short summers of the glacial period. Thus the very existence of such an animal in such remote localities closely associated with those implying almost an arctic winter climate appears to afford a strong support to the argument for the existence of warm inter-glacial or post-glacial periods.

Evidence of Interglacial Warm Periods on the Continent and in North America.—Besides the evidence already adduced from our own islands, many similar facts have been noted in other countries. In Switzerland two glacial periods are distinctly recognised, between which was a warm period when vegetation was so luxuriant as to form beds of lignite sufficiently thick to be worked for coal. The plants found in these deposits are similar to those now inhabiting Switzerland—pines, oaks, birches, larch, etc., but numerous animal remains are also found, showing that the country was then inhabited by an elephant (Elephas antiquus), a rhinoceros (Rhinoceros megarhinus), the urus (Bos primigenius), the red deer (Cervus elephas), and the cave-bear, (Ursus spelœus); and there were also abundance of insects.42

In Sweden also there are two "tills," the lower one having been in places partly broken up and denuded before the upper one was deposited, but no interglacial deposits have yet been found. In North America more complete evidence has been obtained. On the shores of Lake Ontario sections are exposed showing three separate beds of "till" with intervening stratified deposits, the lower one of which has yielded many plant remains and fresh-water organisms. These deposits are seen to extend continuously for more than nine miles, and the fossiliferous interglacial beds attain a thickness of 140 feet. Similar beds have been discovered near Cleveland, Ohio, consisting, first of "till" at the lake-level, secondly of about 48 feet of sand and loam, and thirdly of unstratified "till" full of striated stones—six feet thick.43 On the other side of the continent, in British Columbia, Mr. G. M. Dawson, geologist to the North American Boundary Commission, has discovered similar evidence of two glaciations divided from each other by a warm period.

This remarkable series of observations, spread over so wide an area, seems to afford ample proof that the glacial epoch did not consist merely of one process of change, from a temperate to a cold and arctic climate, which having reached a maximum, then passed slowly and completely away; but that there were certainly two, and probably several more alternations of arctic and temperate climates.

It is evident, however, that if there have been, not two only, but a series of such alternations of climate, we could not possibly expect to find more than the most slender indications of them, because each succeeding ice-sheet would necessarily grind down or otherwise destroy much of the superficial deposits left by its predecessors, while the torrents that must always have accompanied the melting of these huge masses of ice would wash away even such fragments as might have escaped the ice itself. It is a fortunate thing therefore, that we should find any fragments of these interglacial deposits containing animal and vegetable remains; and just as we should expect, the evidence they afford seems to show that the later phase of the cold period was less severe than the earlier. Of such deposits as were formed on land during the coming on of the glacial epoch when it was continually increasing in severity hardly a trace has been preserved, because each succeeding extension of the ice being greater and thicker than the last, destroyed what had gone before it till the maximum was reached.

Migrations and Extinction of Organisms caused by the Glacial Epoch.—Our last glacial epoch was accompanied by at least two considerable submergences and elevations of the land, and there is some reason to think, as we have already explained, that the two classes of phenomena are connected as cause and effect. We can easily see how such repeated submergences and elevations would increase and aggravate the migrations and extinctions that a glacial epoch is calculated to produce. We can therefore hardly fail to be right in attributing the wonderful changes in animal and vegetable life that have occurred in Europe and N. America between the Miocene Period and the present day, in part at least, to the two or more cold epochs that have probably intervened. These changes consist, first, in the extinction of a whole host of the higher animal forms, and secondly, in a complete change of types due to extinction and migration, leading to a much greater difference between the vegetable and animal forms of the eastern and western hemisphere than before existed. Many large and powerful mammalia lived in our own country in Pliocene times and apparently survived a part of the glacial epoch; but when it finally passed away they too had disappeared, some having become altogether extinct while others continued to exist in more southern lands. Among the first class are the sabre-toothed tiger, the extinct Siberian camel (Merycotherium), three species of elephant, two of rhinoceros, two bears, five species of deer, and the gigantic beaver; among the latter are the hyæna, bear, and lion, which are considered to be only varieties of those which once inhabited Britain. Down to Pliocene times the flora of Europe was very similar to that which now prevails in Eastern Asia and Eastern North America. The late Professor Asa Gray has pointed out that hundreds of species of trees and shrubs of peculiar genera which still flourish in those countries are now completely wanting in Europe, and there is good reason to believe that these were exterminated during the glacial period, being cut off from a southern migration, first by the Alps, and then by the Mediterranean; whereas in eastern America and Asia the mountain chains run in a north and south direction, and there is nothing to prevent the flora from having been preserved by a southward migration into a milder region.44

Our next two chapters will be devoted to a discussion of the causes which brought about the glacial epoch, and that still more extraordinary climatic phenomenon—the mild climate and luxuriant vegetation of the Arctic zone. If my readers will follow me with the care and attention so difficult and interesting a problem requires and deserves, they will find that I have grappled with all the more important facts which have to be accounted for, and have offered what I believe is the first complete and sufficient explanation of them. The important influence of climatal changes on the dispersal of animals and plants is a sufficient justification for introducing such a discussion into the present volume.

CHAPTER VIII

THE CAUSES OF GLACIAL EPOCHS

Various Suggested Causes—Astronomical Causes of Changes of Climate—Difference of Temperature caused by Varying Distance of the Sun—Properties of Air and Water, Snow and Ice, in Relation to Climate—Effects of Snow on Climate—High Land and Great Moisture Essential to the Initiation of a Glacial Epoch—Perpetual Snow nowhere Exists on Lowlands—Conditions Determining the Presence or Absence of Perpetual Snow—Efficiency of Astronomical Causes in Producing Glaciation—Action of Meteorological causes in Intensifying Glaciation—Summary of Causes of Glaciation—Effect of Clouds and Fog in cutting off the Sun's Heat—South Temperate America as Illustrating the Influence of Astronomical Causes on Climate—Geographical Changes how far a Cause of Glaciation—Land acting as a Barrier to Ocean-currents—The theory of Interglacial Periods and their Probable Character—Probable Effect of Winter in Aphelion on the Climate of Britain—The Essential Principle of Climatal Change Restated—Probable Date of the last Glacial Epoch—Changes of the Sea-level dependent on Glaciation—The Planet Mars as bearing on the Theory of Excentricity as a Cause of Glacial Epochs.

No less than seven different causes have been at various times advanced to account for the glacial epoch and other changes of climate which the geological record proves to have taken place. These, as enumerated by Mr. Searles V. Wood, Jun., are as follows:—

1. A decrease in the original heat of our planet.

2. Changes in the obliquity of the ecliptic.

3. The combined effect of the precession of the equinoxes and of the excentricity of the earth's orbit.

4. Changes in the distribution of land and water.

5. Changes in the position of the earth's axis of rotation.

6. A variation in the amount of heat radiated by the sun.

7. A variation in the temperature of space.

Of the above, causes (1) and (2) are undoubted realities; but it is now generally admitted that they are utterly inadequate to produce the observed effects. Causes (5) (6) and (7) are all purely hypothetical, for though such changes may have occurred there is no evidence that they have occurred during geological time; and it is besides certain that they would not, either singly or combined, be adequate to explain the whole of the phenomena. There remain causes (3) and (4), which have the advantage of being demonstrated facts, and which are universally admitted to be capable of producing some effect of the nature required, the only question being whether, either alone or in combination, they are adequate to produce all the observed effects. It is therefore to these two causes that we shall confine our inquiry, taking first those astronomical causes whose complex and wide reaching effects have been so admirably explained and discussed by Dr. Croll in numerous papers and in his work—"Climate and Time in their Geological Relations."

DIAGRAM SHOWING THE ALTERED POSITION OF THE POLES AT INTERVALS OF 10,500 YEARS PRODUCED BY THE PRECESSION OF THE EQUINOXES AND THE MOTION OF THE APHELION; AND ITS EFFECT ON CLIMATE DURING A PERIOD OF HIGH EXCENTRICITY.

Astronomical Causes of Changes of Climate.—The earth moves in an elliptical orbit round the sun, which is situated in one of the foci of the ellipse, so that the distance of the sun from us varies during the year to a considerable amount. Strange to say we are now three millions of miles nearer to the sun in winter than in summer, while the reverse is the case in the southern hemisphere; and this must have some effect in making our northern winters less severe than those of the south temperate zone. But the earth moves more rapidly in that part of its orbit which is nearer to the sun, so that our winter is not only milder, but several days shorter, than that of the southern hemisphere. The distribution of land and sea and other local causes prevent us from making any accurate estimate of the effects due to these differences; but there can be no doubt that if our winter were as long as our summer is now and we were also three million miles further from the sun at the former period, a very decided difference of climate would result—our winter would be colder and longer, our summer hotter and shorter. Now there is a combination of astronomical revolutions (the precession of the equinoxes and the motion of the aphelion) which actually brings this change about every 10,500 years, so that after this interval the condition of the two hemispheres is reversed as regards nearness to the sun in summer, and comparative duration of summer and winter; and this change has been going on throughout all geological periods. (See Diagram.) The influence of the present phase of precession is perhaps seen in the great extension of the antarctic ice-fields, and the existence of glaciers at the sea-level in the southern hemisphere, in latitudes corresponding to that of England; but it is not supposed that similar effects were produced with us at the last cold period, 10,500 years ago, because we are exceptionally favoured, by the Gulf-stream warming the whole North Atlantic ocean and by the prevalence of westerly winds which convey that warmth to our shores; and also by the comparatively small quantity of high land around the North Pole which does not encourage great accumulations of ice. But besides this change in the relation of our seasons to the earth's aphelion and perihelion there is another and still more important astronomical factor in the change of magnitude of the excentricity itself. This varies very largely, though very slowly, and it is now nearly at a minimum. It also varies very irregularly; but its amount has been calculated for several million years back. Fifty thousand years ago it was rather less than it is now, but it then increased, and when we come to a hundred thousand years ago there is a difference of eight and a half millions of miles between our distance from the sun in aphelion and perihelion (as the most distant and nearest points of the earth's orbit are termed). At a hundred and fifty thousand years back it had decreased somewhat—to six millions of miles; but then it increased again, till at two hundred thousand years ago it was ten and a quarter, and at two hundred and ten thousand years ten and a half millions of miles. By reference to the accompanying diagram, which includes the last great period of excentricity, we find, that for the immense period of a hundred and sixty thousand years (commencing about eighty thousand years ago) the excentricity was very great, reaching a maximum of three and a half times its present amount at almost the remotest part of this period, at which time the length of summer in one hemisphere and of winter in the other would be nearly twenty-eight days in excess. Now, during all this time, our position would change, as above described (and as indicated on the diagram), every ten thousand five hundred years; so that we should have alternate periods of very long and cold winters with short hot summers, and short mild winters with long cool summers. In order to understand the important effects which this would produce we must ascertain two things—first, what actual difference of temperature would be caused by varying distances of the sun, and, secondly, what are the properties of snow and ice in regard to climate.

DIAGRAM OF EXCENTRICITY AND PRECESSION.

The dark and light bands mark the phases of precession, the dark showing short mild winters, and the light long cold winters, the contrast being greater as the excentricity is higher. The horizontal dotted line shows the amount of the present excentricity. The figures show the maxima and minima of excentricity during the last 300,000 years from Dr. Croll's Tables.

Differences of Temperature Caused by Varying Distances of the Sun.—On this subject comparatively few persons have correct ideas owing to the unscientific manner in which we reckon heat by our thermometers. The zero of Fahrenheit's thermometer is thirty-two degrees below the freezing point of water, and that of the centigrade thermometer, the freezing point itself, both of which are equally misleading when applied to cosmical problems. If we say that the mean temperature of a place is 50° F., or 10° C., these figures tell us nothing of how much the sun warms that place, because if the sun were withdrawn the temperature would fall far below either of the zero points. In the last Arctic Expedition a temperature of -74° F. was registered, or 106° below the freezing point of water; and as at the same time the earth, at a depth of two feet, was only, -13° F. and the sea water +28° F., both influencing the temperature of the air, we may be sure that even this intense cold was not near the possible minimum temperature. By various calculations and experiments which cannot be entered upon here, it has been determined that the temperature of space, independent of solar (but not of stellar) influence, is about -239° F., and physicists almost universally adopt this quantity in all estimates of cosmical temperature. It follows, that if the mean temperature of the earth's surface at any time is 50° F. it is really warmed by the sun to an amount measured by 50 + 239 = 289° F., which is hence termed its absolute temperature. Now during the time of the glacial epoch the greatest distance of the sun in winter was 98¼ millions of miles, whereas it is now, in winter, only 91½ millions of miles, the mean distance being taken as 93 million miles. But the quantity of heat received from the sun is inversely as the square of the distance, so that it would then be in the proportion of 8,372 to 9,613 now, or nearly one seventh less than its present amount. The mean temperature of England in January is about 37° F., which equals 276° F. of absolute temperature. But the above-named fraction of 276° is 237, the difference, 39, representing the amount which must be deducted to obtain the January temperature during the glacial epoch, which will therefore be -2° F. But this is a purely theoretic result. The actual temperature at that time might have been very different from this, because the temperature of a place does not depend so much on the amount of heat it receives directly from the sun, as on the amount brought to it or carried away from it by warm or cold winds. We often have it bitterly cold in the middle of May when we are receiving as much sun heat as many parts of the tropics, but we get cold winds from the iceberg-laden North Atlantic, and this largely neutralises the effect of the sun. So we often have it very mild in December if south-westerly winds bring us warm moist air from the Gulf-stream. But though the above method does not give correct results for any one time or place, it will be more nearly correct for very large areas, because all the sensible surface-heat which produces climates necessarily comes from the sun, and its proportionate amount may be very nearly calculated in the manner above described. We may therefore say, generally, that during our winter, at the time of the glacial epoch, the northern hemisphere was receiving so much less heat from the sun as was calculated to lower its surface temperature on an average about 39° F., while during the height of summer of the same period it would be receiving so much more heat as would suffice, other conditions being equal, to raise its mean temperature about 48° above what it is now. The winter, moreover, would be long and the summer short, the difference being twenty-six days.

We have here certainly an amount of cold in winter amply sufficient to produce a glacial period,45 especially as this cold would be long continued; but at the same time we should have almost tropical heat in summer, although that season would be somewhat shorter. How then, it may be asked, could such a climate have the effect supposed? Would not the snow that fell in winter be all melted by the excessively hot summer? In order to answer this question we must take account of certain properties of water and air, snow and ice, to which due weight has not been given by writers on this subject.

Properties of Air and Water, Snow and Ice, in Relation to Climate.—The great aerial ocean which surrounds us has the wonderful property of allowing the heat-rays from the sun to pass through it without its being warmed by them; but when the earth is heated the air gets warmed by contact with it, and also to a considerable extent by the heat radiated from the warm earth, because, although pure dry air allows such dark heat-rays to pass freely, yet the aqueous vapour and carbonic acid in the air intercept and absorb them. But the air thus warmed by the earth is in continual motion owing to changes of density. It rises up and flows off, owing to the greater weight of the cooler air which forces it up and takes its place; and thus heat can never accumulate in the atmosphere beyond a very moderate degree, the excessive sun-heat of the tropics being much of it carried away to the upper atmosphere and radiated into space. Water also is very mobile; and although it receives and stores up a great deal of heat, it is for ever dispersing it over the earth. The rain which brings down a certain portion of heat from the atmosphere, and which often absorbs heat from the earth on which it falls, flows away in streams to the ocean; while the ocean itself, constantly impelled by the winds, forms great currents, which carry off the surplus heated water of the tropics to the temperate and even to the polar regions, while colder water flows from the poles to ameliorate the heat of the tropics. An immense quantity of sun-heat is also used up in evaporating water, and the vapour thus produced is conveyed by the aerial currents to distant countries, where, on being condensed into rain, it gives up much of this heat to the earth and atmosphere.

The power of water in carrying away heat is well exhibited by the fact of the abnormally high temperature of arid deserts and of very dry countries generally; while the still more powerful influence of moving air may be appreciated, by considering the effects of even our northern sun in heating a tightly-closed glass house to far above the temperature produced by the vertical sun of the equator where the free air and abundance of moisture exert their beneficial influence. Were it not for the large proportion of the sun's heat carried away by air and water the tropics would become uninhabitable furnaces—as would indeed any part of the earth where the sun shone brightly throughout a summer's day.

We see, therefore, that the excess of heat derived from the sun at any place cannot be stored up to an important amount owing to the wonderful dispersing agency of air and water; and though some heat does penetrate the ground and is stored up there, this is so little in proportion to the whole amount received, and the larger part of it is so soon given out from the surface layers, that any surplus heat that may be thus preserved during one summer of the temperate zones rarely or never remains in sufficient quantity to affect the temperature of the succeeding summer, so that there is no such thing as an accumulation of earth-heat from year to year. But, though heat cannot, cold can be stored up to an almost unlimited amount, owing to the peculiar property water possesses of becoming solid at a moderately low temperature; and as this is a subject of the very greatest importance to our inquiry—the whole question of the possibility of glacial epochs and warm periods depending on it—we must consider it in some detail.

Effects of Snow on Climate.—Let us then examine the very different effects produced by water falling as a liquid in the form of rain, or as a solid in the form of snow, although the two may not differ from each other more than two or three degrees in temperature. The rain, however much of it may fall, runs off rapidly into streams and rivers, and soon reaches the ocean, a small portion only sinking into the earth and another portion evaporating into the atmosphere. If cold it cools the air and the earth somewhat while passing through or over them, but produces no permanent effect on temperature, because a few hours of sunshine restore to the air or the surface-soil all the heat they had lost. But if snow falls for a long time, the effect, as we all know, is very different, because it has no mobility. It remains where it fell and becomes compacted into a mass, and it then keeps the earth below it and the air above, at or near the freezing-point till it is all melted. If the quantity is great it may take days or weeks to melt; and if snow continues falling it goes on accumulating all over the surface of a country (which water cannot do), and may thus form such a mass that the warmth of the whole succeeding summer may not be able to melt it. It then produces perpetual snow, such as we find above a certain altitude on all the great mountains of the globe; and when this takes place cold is rendered permanent, no amount of sun-heat warming the air or the earth much above the freezing-point. This is illustrated by the often-quoted fact that, at 80° N. Lat., Captain Scoresby had the pitch melted on one side of his ship by the heat of the sun, while water was freezing on the other side owing to the coldness of the air.