Studies in the Theory of Descent, Volume II

Now if this wide deviation in every part were in itself no obstacle to the assumption of a designing and re-modelling power, it would become so by the circumstance that all the parts of the organism must stand in the most precise relation to the external conditions of life, if the organism is to be capable of existing – all the parts must be exactly adapted to certain conditions of life. How can this be brought about by a transforming force acting spasmodically? Von Hartmann – who, in spite of his clear perception and widely extended scientific knowledge, cannot possibly possess a strong conviction of that harmony between structure and life-conditions prevailing throughout the whole system of the organism, and which personal research and contemplation are alone able to give – simply bridges over the difficulty by permitting natural selection to come to his aid as an “auxiliary principle” of the re-modelling power. It would not be supposed that naturalists would resort to the same device – nevertheless those who support the phyletic force and per saltum development generally invoke natural selection as the principle which governs adaptation. But when does this agency come into operation? When by germinal metamorphosis a new form has arisen, this, from the first moment of its existence, must be adapted to the new conditions of life or it must perish. No time is allowed for it to continue in an unadapted state throughout a series of generations until adaptation is luckily reached through natural selection. Let us have either natural selection or a phyletic force – both together are inconceivable. If there exists a phyletic force, then it must itself bring about adaptation.

It might perhaps be here suggested that the same objection applies to that process of modification which is effected by small steps, but that it does so only when the change occurs suddenly. This, however, as I have already attempted to show, but very rarely takes place; in many cases (mimicry) the conditions even change in the first place through the change in form and therefore, as is evident, as gradually as the latter. It must be the same in all other cases where transformation of the existing form and not merely extinction of the species concerned takes place. The transmutation must always keep pace with the change in the conditions of life, since if the latter change more rapidly the species could not compete with rival species – it would become extinct.

The abrupt transformation of species implies sudden change in the conditions of life, since a Medusa does not live like a Polype, nor a Trematode like its “nurse.” For this reason it is impossible that natural selection can be an aiding principle of “heterogeneous generation.” If such abrupt transformation takes place it must produce the new form instantly equipped for the struggle for existence, and adapted in all its organs and systems of organs to the special conditions of its new life. But would not this be “pure magic”? It is not thereby even taken into consideration that here – as in the cases of mimicry – time and place must agree. The requirements of a pre-established harmony (“prästabilirte Harmonie”) further demand that an animal fitted for special conditions of life should only make its appearance at that precise period of the earth’s history when these special conditions are all fulfilled, and so forth.

But he who has learnt to perceive the numerous and fine relations which, in every species of animal, bring the details of structure into harmony with function, and who keeps in view the impelling power of these conditions, cannot possibly hold to the idea of a per saltum development of animal forms. If development has taken place, it must have occurred gradually and by minute steps – in such a manner indeed that each modification had time to become equilibrated to the other parts, and in this way a succession of modifications gradually brought about the total transformation of the organism, and at the same time secured complete adaptation to new conditions of life.

Not only abrupt modification however, but every transformation is to be rejected when based upon the interference of a metaphysical principle of development. Those to whom the arguments already advanced against such a principle appear insufficient may once more be asked, how and where should this principle properly interfere? I am of opinion that one effect can have but one sufficient cause; if this suffices to produce it, no second cause is required. The hand of a watch necessarily turns once round in a circle in a given time as soon as the spring which sets the mechanism in movement is wound up; in an unwound watch a skilful finger can perhaps give the same movement to the hand, but it is impossible that the latter can receive both from the operator and from the spring at the same time, the same motion as that which it would receive through either of these two powers alone. In the same manner it appears to me that the variations which lead to transformation cannot be at the same time determined by physical and by metaphysical causes, but must depend upon either one or the other.

On no side will it be disputed that at least one portion of the processes of organic life depends upon the mechanical co-operation of physical forces. How is it conceivable that sudden pauses should occur in the course of these causal forces, and that a directive power should be substituted therefor, the latter subsequently making way again for the physical forces? To me this is as inconceivable as the idea that lightning is the electric discharge of a thunder-cloud, of which the formation and electrical tension depends upon causal forces, and of which the time and place are purely determined by such forces, but that Jupiter has it nevertheless in his power to direct the lightning flash according to his will on to the head of the guilty.

Now although I deny the possibility or conceivability of the contemporaneous co-operation of teleological and of causal forces in producing any effect, and although I maintain that a purely mechanical conception of the processes of nature is alone justifiable, I nevertheless believe that there is no occasion for this reason to renounce the existence of, or to disown, a directive power; only we must not imagine this to interfere directly in the mechanism of the universe, but to be rather behind the latter as the final cause of this mechanism.

Von Baer himself points this out to us, although he does not follow up the complete consequences of his arguments. He especially insists in his book, which abounds in beautiful and grand ideas, that the notions of necessity (causality) and of purpose by no means necessarily exclude one another, but rather that they can be connected together in a certain manner. Thus, the watchmaker attains his end, the watch, by combining the elastic force of a spring with wheel-work, i. e. by utilizing physical necessities; the farmer accomplishes his purpose, that of obtaining a crop of corn, by sowing the seed in suitable land, but the seed must germinate as an absolute necessity when exposed to the influences of warmth, soil, moisture, &c. Thus, in these instances a chain of necessities is undoubtedly connected with a teleological force, the human will; and it directly follows from such cases that wherever we see an aim or result attained through necessities, the directive force does not interrupt the course of the series of necessities which have already commenced, but is active before the first commencement of these necessities, since it combines and sets the latter in movement. From the moment when the mechanism of the watch is combined harmoniously and the spring wound up, it goes without the further interference of the watchmaker, just as the corn-seed when once placed in the earth develops into a plant without assistance from the farmer.

If we apply this argument to the development of the organic world, those who defend mechanical development will not be compelled to deny a teleological power, only they would have with Kant139 to think of the latter in the only way in which it can be conceived, viz. as a Final Cause.

In the region of inorganic nature nobody any longer doubts the purely mechanical connection of the phenomena. Sunshine and rain do not now appear to us to be whims of a deity, but divine natural laws. As the knowledge of the processes of nature advances, the point where the divine power designedly interrupts these processes must be removed further back; or, as the author of the criticism of the philosophy of the Unconscious140 expresses it, all advance in the knowledge of natural processes depends “upon the continual elimination of the idea of the miraculous.” We now believe that organic nature must be conceived as mechanical. But does it thereby follow that we must totally deny a final Universal Cause? Certainly not; it would be a great delusion if any one were to believe that he had arrived at a comprehension of the universe by tracing the phenomena of nature to mechanical principles. He would thereby forget that the assumption of eternal matter with its eternal laws by no means satisfies our intellectual need for causality. We require before everything an explanation of the fact that relationships everywhere exist between the parts of the universe – that atoms everywhere act upon one another.141 He who can content himself with the assumption of matter may do so, but he will not be able to show that the assumption of a Universal Cause underlying the laws of nature is erroneous.

It will not be said that there is no advantage in assuming such a Final Cause, because we cannot conceive it, and indeed cannot so much as demonstrate it with certainty. It certainly lies beyond our power of conception, in the obscure region of metaphysics, and all attempts to approach it have never led to anything but an image or a formula. Nevertheless there is an advance in knowledge in the assumption of this Cause which well admits of comparison with those advances which have been led to by certain results of the new physiology of the senses. We now know that the images which give us our sense of the external world are not “actual representations having any degree of resemblance,”142 but are only signs for certain qualities of the outer world, which do not exist as such in the latter, but belong entirely to our consciousness. Thus we know for certain that the world is not as we perceive it – that we cannot perceive “things in their essence” – and that the reality will always remain transcendental to us. But who will deny that in this knowledge there is a considerable advance, in spite of its being for the most part of a negative character? But just as we must assume behind the phenomenal world of our senses an actual world of the true nature of which we receive only an incomplete knowledge (i. e. a knowledge corresponding only in reality with the relations of time and space), so behind the co-operating forces of nature which “aim at a purpose” must we admit a Cause, which is no less inconceivable in its nature, and of which we can only say one thing with certainty, viz., that it must be teleological. Just as the former first leads us to perceive the true value of our sensual impressions, so does the latter knowledge lead us to foresee the true significance of the mechanism of the universe.

It is true that in neither case do we learn more than that there is something present which we do not perceive, but in both instances this knowledge is of the greatest value. The consciousness that behind that mechanism of the universe which is alone comprehensible to us there still lies an incomprehensible teleological Universal Cause, necessitates quite a different conception of the universe – a conception absolutely opposed to that of the materialist. Most correctly and beautifully does Von Baer say that “a purpose cannot be otherwise conceived by us than as proceeding from a will and consciousness; in this would the ‘aiming at a purpose,’ which appears to us as reasonable as it is necessary, have its deepest root.” If we conceive in this world a divine Universal Power exercising volition as the ultimate basis of matter and of the natural laws resident therein, we thus reconcile the apparent contradiction between the mechanical conception and teleology. In the same way that Von Hartmann, somewhere speaks of the immanent teleology of a machine, we might speak of the immanent teleology of the universe, because the single forces of matter are so exactly adjusted that they must give rise to the projected world, just as the wheels and levers of a machine bring forth a required manufactured article. I admit that these are grossly anthropomorphic ideas. But as mortals can we have any other ideas? Is not the notion of purpose in itself an equally anthropomorphic one? and is there any certainty that the idea of causality is less so? Do we know that causality is unlimited, or that it is universally valid? In the absence of this knowledge, should it not be permissible to satisfy as far as we can the craving of the human mind for a spiritual First Cause of the universe, by speaking of it in terms conceivable to human understanding? We can take up such a final position and still be conscious that we thereby form no certain conception, and indeed come no nearer to the reality. The materialist still makes use of the notion of “eternity,” and frequently handles it as though it were a perfectly known quantity. We nevertheless do not seriously believe that by the expression “eternal matter,” any true idea resulting from human experience is gained.

If it is asked, however, how that which in ourselves and in the remainder of the animal world is intellectual and perceptive, which thinks and wills, is ascribable to a mechanical process of organic development – whether the development of the mind can be conceived as resulting from purely mechanical laws? I answer unhesitatingly in the affirmative with the pure materialist, although I do not agree with him as to the manner in which he derives these phenomena from matter, since thinking and extension are heterogeneous things, and one cannot be considered as a product of the other. But why should not the ancient notion of “conscious matter” given out by Maupertuis and Robinet, not be again entertained, as pointed out in recent times by Fechner?143 Would there not thus be found a useful formula for explaining phenomena hitherto quite incomprehensible?

Von Hartmann in criticizing himself, designates the sensibility of atoms as an “almost inevitable hypothesis” (p. 62), “inevitable because if sensibility were not a general and original property of the constituent elements of matter, it would be absolutely incomprehensible how through its potentiality and integration that sensibility known to us as being possessed by the organism could have arisen.” “It is impossible that from purely external elements devoid of all internality (Innerlichkeit) there should suddenly appear, by a certain mode of combination, an internality which becomes more and more richly developed. The more certainly science becomes convinced that in the sphere of externality (Äusserlichkeit) the higher (organic) phenomena are only results of combination, or are the aggregate phenomena of the elementary atomic forces, the more surely, when she once seriously concerns herself with this other question, will she not fail to be convinced that the sensibility possessed by higher stages of consciousness can be only combination-results, or the aggregate phenomena of the elementary sensations of atoms, although these atomic sensations as such always remain below the level of the higher combinations of consciousness.” In confusing this double-sided nature of the objective phenomenon “lies the main error of all materialism and of all subjective idealism. Just as the attempt of the latter (subjective idealism) to construct the external phenomena of existence in space out of functions of internality and their combinations is impossible, so is the endeavour of the former (materialism) to build up internal sensation out of any combinations of force acting externally in space equally impossible.”

I have no intention of going any deeper into these questions. I mention them only in order to point out that even from this side there appears to me no obstacle in the way of a purely mechanical conception of the processes of the universe. The naturalist may be excused if he attempts to penetrate into the region of philosophy; it arises from the wish to be able to contribute a little towards the reconciliation of the latest knowledge of the naturalist with the religious wants of the human mind – towards the aim striven for by both sides, viz. a satisfactory and harmonious view of the universe, according with the state of knowledge of our time.

I believe that I have shown that the theory of selection by no means leads – as is always assumed – to the denial of a teleological Universal Cause and to materialism, and I thereby hope that I have cleared the way for this doctrine, the importance of which it is scarcely possible to over-estimate. Many, and not the most ill-informed, do not get so far as to make an unbiassed examination into the facts, because they are at the outset alarmed by the to them inevitable consequence of the materialistic conception of the universe. Mechanism and teleology do not exclude one another, they are rather in mutual agreement. Without teleology there would be no mechanism, but only a confusion of crude forces; and without mechanism there would be no teleology, for how could the latter otherwise effect its purpose?144

Von Hartmann correctly says: – “The most complete mechanism conceivable is likewise the most completely conceivable teleology.” We may thus represent the phenomenal universe as such a completely conceivable mechanism. With this conception vanish all apprehensions that the new views would cause man to lose the best that he possesses – morality and purely human spiritual culture. He who, with Von Baer, considers the laws of nature as the “permanent expressions of the will of a creative principle,” will clearly perceive that a further advance in the knowledge of these laws need not divert man from the path of increasing improvement, but must further him in this course – that the knowledge of truth, whatever may be its purport, cannot possibly be considered a backward step. Let us take our stand boldly on the ground of new knowledge, and accept the direct consequences thereof, and we shall not be obliged to give up either morality or the comforting conviction of being part of an harmonious world, as a necessary member capable of development and perfection.

Any other mode of interference by a directive teleological power in the processes of the universe than by the appointment of the forces producing them, is however, at least to the naturalist, inadmissible. We are still far removed from completely understanding the mechanism by means of which the organic world is evoked – we still find ourselves at the very beginning of knowledge. We are, however, already convinced that both the organic and the inorganic worlds are dependent only upon mechanical forces, for to this conclusion we are led, not only by the results of investigators who have restricted themselves to limited provinces, but also by the most general considerations. But although the force of these arguments may not be acknowledged, and although one might maintain that the inductional proofs against the existence of a “phyletic vital force” have been directed only against points of detail, or have never been completely demonstrated, i. e. for all points, it must nevertheless be conceded, that for the naturalist the mechanical conception of Nature is the only one possible – that he is not at all justified in abandoning this view so long as the interference of teleological forces in the course of the processes of organic development has not been demonstrated to him. Thus, it will not be immaterial whether a conception of Nature which to many seems inevitable is consistent with the idea of universal design, or a final directive universal principle, since the value which we may attach to our own lives and aims, essentially depends thereon. The final and main result of this essay will thus be found in the attempted demonstration that the mechanical conception of Nature very well admits of being united with a teleological conception of the Universe.

THE END

ERRATA

Page 81, line 8 from top, and throughout essay, for “Daphnidæ” read “Daphniidæ.”

Page 95, line 3 from bottom, for “Daphnoidea” read “Daphniacea.”

Page 166, line 7 from bottom (note), for “p. 438” read “p. 433.”

Page 245, line 17 from bottom (note), for “Ställ” read “Stoll.”

Page 263, after the word “insects” (bottom line of note), add, “but the whole marking is suggestive of distastefulness.”

Page 296, line 3 from bottom, for “Stähelina – collector” read “Stähelin – a collector.”

Page 305, line 5 from bottom (note), for “In 1869” read “In 1865.”

Page 434, bottom line of note, for “Geometræ” read “Bombycidæ.”

Page 494, line 2 from top, for “from which a larval form” read “from a larval form which.”

Page 542, line 12 from top, for “Dione Vanilla” read “Dione Vanillæ.”

Page 544, line 15 from bottom, for “Siderome” read “Siderone.”

1

[The slight variability in the colour of this pupa, opens up the interesting question of the photographic sensitiveness of this and other species, which is stated to cause them to assimilate in colour to the surface on which the larva undergoes its final ecdysis. Some experiments upon this subject have been recorded by Mr. T. W. Wood, Proc. Ent. Soc. 1867, p. xcix, but the field is still almost unexplored. R.M.]

2

“Über den Einfluss der Isolirung auf die Artbildung.” Leipzig, 1872, p. 20.

3

In some instances Deilephila Lineata has also been seen by day hovering over flowers.

4

It is true that I only reared one brood, but from this fifty specimens were obtained. It would be interesting to know whether this variety of the caterpillar is distributed over the whole of Southern Europe.

5

In this sense Lubbock says: – “It is evident that creatures which, like the majority of insects, live during the successive periods of their existence in very different circumstances, may undergo considerable changes in their larval organization in consequence of forces acting on them while in that condition; not, indeed, without affecting, but certainly without affecting to any corresponding extent, their ultimate form.” – “Origin and Metamorphoses of Insects,” London, 1874, p. 39.

6

“Grundzüge der Zoologie,” 1875.

7

[Lepidopterists are of course aware that even these distinctions are not absolute, as no single character can be named which does not also appear in certain moths. The definition in this case, as in that of most other groups of animals and plants, is only a general one. See, for instance, Westwood’s “Introduction to the Classification of Insects,” vol. ii. pp. 330–332. Also some remarks by C. V. Riley in his “Eighth Annual Report” on the insects of Missouri, 1876, p. 170. With reference to the antennæ as a distinguishing character, see Mr. A. G. Butler’s article in “Science for All,” 1880, part xxvii. p. 65. R.M.]

8

The genus of Morphinæ, Discophora, possesses hairs very similar to those of the genus Cnethocampa belonging to the Bombycidæ.

9

[The larvæ of genera 14, Phyciodes, and 35, Crenis, are likewise spiny. See Edwards’ “Butt. of N. Amer.” vol. ii. for figures of the caterpillar of Phyc. Tharos: for notes on the larvæ of Crenis Natalensis and C. Boisduvali see a paper by W. D. Gooch, “Entomologist,” vol. xiv. p. 36. The larvæ of genus 55, Ageronia, are also spiny. (See Burmeister’s figure of A. Arethusa, “Lép. Rép. Arg.” Pl. V. Fig. 4). The larvæ of genus 98, Aganisthos, also appear to be somewhat spiny (see Burmeister’s figure of A. Orion, loc. cit. Pl. V. Fig. 6), and this raises the question as to whether the genus is correctly located in its present position. The larvæ of the following genera figured in Moore’s “Lepidoptera of Ceylon,” parts i. and ii., are all spiny: – 6, Cirrochroa (Pl. XXXII.); 7, Cynthia (Pl. XXVI.); 27, Kallima (Pl. XIX.); and 74, Parthenos (Pl. XXIV.). Many species of caterpillars which are spiny when adult appear to be spineless, or only slightly hairy when young. See Edwards’ figures of Melitæa Phaeton, Argynnis Diana, and Phyc. Tharos (loc. cit.) and his description of the larva of Arg. Cybele, “Canad. Entom.” vol. xii. p. 141. The spiny covering thus appears to be a character acquired at a comparatively recent period in the phyletic development. R.M.]